Cardamine bulbosa |
Cardamine occidentalis |
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bulb bittercress, bulbous bitter-cress, spring cress |
big western bitter-cress, western bitter-cress, western toothwort |
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Habit | Perennials; glabrous or sparsely pubescent distally. | Perennials; glabrous or hirsute. |
Rhizomes | (tuberous at stem base, sometimes also at intervals) subglobose, lobed, 4–15 mm diam. (fleshy). |
(tuberiform, fragile), ovoid or globose at base of stem, 3–10 mm diam., (fleshy). |
Stems | erect, unbranched, (1–)2–6 dm, glabrous or sparsely pubescent on distal 1/2 (trichomes 0.02–0.1 mm). |
(simple from base), erect to ascending, (not flexuous), unbranched or branched distally, 1–5 dm, glabrous or pubescent proximally. |
Basal leaves | not rosulate, pinnately compound, (3 or) 5 (or 7)-foliolate, 2–10 cm, leaflets petiolulate or subsessile; petiole 0.5–6.5 cm; lateral leaflets petiolulate or subsessile, blade similar to terminal, ovate, smaller, margins entire; terminal leaflet (petiolule 0.03–0.18 cm), blade orbicular to broadly ovate or subcordate, 0.5–2 cm × 7–25 mm, base cordate to rounded, margins entire or repand, (surfaces glabrous). |
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Rhizomal leaves | simple, (2–)4–13(–16) cm; petiole (1.5–)2.5–10(–13) cm; blade usually reniform to cordate or ovate, rarely oblong, (1–)2–4(–6) cm, base obtuse to cordate, margins usually repand or entire, rarely shallowly dentate. |
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Cauline leaves | (2–)4–10(–14), simple, petiolate or sessile; (middle ones) shortly petiolate or (distally) sessile, base not auriculate; blade ovate to oblong, or oblong-linear to lanceolate, 3–6(–9) cm × 10–30(–45) mm, margins entire, repand, or dentate (margins minutely pubescent). |
3–7, (3 or) 5 or 7-foliolate (middle ones 5 or 7-foliolate, smaller distally, becoming 3-foliolate), petiolate; petiole 0.5–3 cm, base not auriculate; lateral leaflets similar to terminal, smaller; terminal leaflet blade obovate to oblanceolate, 0.5–2.6 cm × 3–13 mm, margins shallowly toothed, entire, or repand. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 2.5–4.5 × 1.5–2 mm, lateral pair not saccate basally, (glabrous); petals usually white, rarely pale pink, obovate, (6–)7–12(–16) × 3–5 mm, (short-clawed, apex rounded); filaments: median pairs 4.5–7 mm, lateral pair 2–3.5 mm; anthers oblong, 1–1.5 mm. |
sepals oblong, 1.7–2 × 1–1.2 mm, lateral pair not saccate basally; petals white, oblanceolate, 4–6 × 1.5–2 mm, (not clawed); filaments: median pairs 2–2.5 mm, lateral pair 1–1.5 mm; anthers ovate, 0.3–0.5 mm. |
Fruiting pedicels | ascending to divaricate, (10–)15–22(–30) mm. |
divaricate-ascending, 7–18 mm. |
Fruits | linear, 2–3.5(–4) cm × 1.4–1.7 mm; ovules 14–24 per ovary; style 2–4(–5) mm. |
linear, (torulose), 1.5–3.3 cm × 1.7–2.2 mm; (valves glabrous or sparsely pubescent); ovules 18–40 per ovary; style 0.5–1.5 mm. |
Seeds | dark orange to greenish yellow, oblong or globose, 1.7–2.1 × 1–1.4 mm. |
brown, ovoid, 1–1.6 × 1–1.2 mm. |
2n | = 16, 56, 64, 80, 96, 112. |
= 64. |
Cardamine bulbosa |
Cardamine occidentalis |
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Phenology | Flowering Mar–Jun. | Flowering Apr–Jul. |
Habitat | Wet grounds, low woodland, moss hummocks, alluvial woods, grassy floodplains, wet pastures, meadows, pinelands, creek bottoms, stream banks, sandy bottoms, ditches, mesic or wet forests, swamps, marshes, seepy bluffs | Muddy grounds, lake margins, shallow streams, meadows |
Elevation | 0-900 m (0-3000 ft) | 150-1500 m (500-4900 ft) |
Distribution |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; SD; TN; TX; VA; VT; WI; WV; MB; ON; QC
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AK; CA; OR; WA; BC
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Source | FNA vol. 7, p. 469. | FNA vol. 7, p. 479. |
Parent taxa | Brassicaceae > tribe Cardamineae > Cardamine | Brassicaceae > tribe Cardamineae > Cardamine |
Sibling taxa | ||
Synonyms | Arabis bulbosa, Arabis rhomboidea, C. rhomboidea, C. rhomboidea var. hirsuta, C. rhomboidea var. parviflora, C. rhomboidea var. pilosa, Dentaria rhomboidea, Dracamine bulbosa | C. pratensis var. occidentalis, C. neglecta |
Name authority | (Schreber ex Muhlenberg) Britton: Prelim. Cat., 4. (1888) | (S. Watson) Howell: Fl. N.W. Amer., 50. (1897) |
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