Cardamine bulbosa |
Cardamine hirsuta |
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bulb bittercress, bulbous bitter-cress, spring cress |
hairy bitter-cress, shotweed |
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Habit | Perennials; glabrous or sparsely pubescent distally. | Annuals; sparsely hirsute basally (at least on petiole of basal leaves), often glabrous distally. |
Rhizomes | (tuberous at stem base, sometimes also at intervals) subglobose, lobed, 4–15 mm diam. (fleshy). |
absent. |
Stems | erect, unbranched, (1–)2–6 dm, glabrous or sparsely pubescent on distal 1/2 (trichomes 0.02–0.1 mm). |
erect, ascending, or decumbent, unbranched or branched basally and/or distally, (0.3–)1–3.5(–4.5) dm, (not flexuous). |
Basal leaves | (persistent to anthesis), rosulate, (5–)8–15(–22)-foliolate, (2–)3.5–15(–17) cm, leaflets petiolulate; petiole 0.5–5 cm, (ciliate); lateral leaflet blade oblong, ovate, obovate, or orbicular, smaller than terminal, margins entire, repand, crenate, or 3-lobed; terminal leaflet (petiolule 0.2–1 cm), blade reniform or orbicular, 0.4–2 cm × 6–30 mm, margins entire, repand, dentate, or 3 or 5-lobed. |
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Rhizomal leaves | simple, (2–)4–13(–16) cm; petiole (1.5–)2.5–10(–13) cm; blade usually reniform to cordate or ovate, rarely oblong, (1–)2–4(–6) cm, base obtuse to cordate, margins usually repand or entire, rarely shallowly dentate. |
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Cauline leaves | (2–)4–10(–14), simple, petiolate or sessile; (middle ones) shortly petiolate or (distally) sessile, base not auriculate; blade ovate to oblong, or oblong-linear to lanceolate, 3–6(–9) cm × 10–30(–45) mm, margins entire, repand, or dentate (margins minutely pubescent). |
1–4(–6), compound as basal, petiolate, [(0.5–)1.2–5.5(–7) cm, including petiole], leaflets petiolulate; blade base not auriculate; leaflets similar to basal. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 2.5–4.5 × 1.5–2 mm, lateral pair not saccate basally, (glabrous); petals usually white, rarely pale pink, obovate, (6–)7–12(–16) × 3–5 mm, (short-clawed, apex rounded); filaments: median pairs 4.5–7 mm, lateral pair 2–3.5 mm; anthers oblong, 1–1.5 mm. |
sepals oblong, 1.5–2.5 × 0.3–0.7 mm, lateral pair not saccate; petals (sometimes absent) white, spatulate, 2.5–4.5(–5) × 0.5–1.1 mm; (stamens usually 4, lateral pair often absent, rarely 5 or 6); filaments 1.8–3 mm; anthers ovate, 0.3–0.5 mm. |
Fruiting pedicels | ascending to divaricate, (10–)15–22(–30) mm. |
erect to ascending, (2–)3–10(–14) mm. |
Fruits | linear, 2–3.5(–4) cm × 1.4–1.7 mm; ovules 14–24 per ovary; style 2–4(–5) mm. |
linear, (torulose), (0.9–)1.5–2.5(–2.8) cm × (0.8–)1–1.4 mm, (often appressed to rachis); ovules 14–40 per ovary; style 0.1–0.6(–1) mm. |
Seeds | dark orange to greenish yellow, oblong or globose, 1.7–2.1 × 1–1.4 mm. |
light brown, oblong or subquadrate, 0.9–1.3(–1.5) × 0.6–0.9(–1.1) mm, (narrowly margined). |
2n | = 16, 56, 64, 80, 96, 112. |
= 16. |
Cardamine bulbosa |
Cardamine hirsuta |
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Phenology | Flowering Mar–Jun. | Flowering Feb–Jul. |
Habitat | Wet grounds, low woodland, moss hummocks, alluvial woods, grassy floodplains, wet pastures, meadows, pinelands, creek bottoms, stream banks, sandy bottoms, ditches, mesic or wet forests, swamps, marshes, seepy bluffs | Roadsides, clearings, disturbed sites, slopes, cedar glades, mixed woods, meadows, fields, waste grounds, damp places, grassy areas |
Elevation | 0-900 m (0-3000 ft) | 0-700 m (0-2300 ft) |
Distribution |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; SD; TN; TX; VA; VT; WI; WV; MB; ON; QC
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AL; AR; CA; CT; DC; DE; FL; GA; IL; KY; LA; MA; MD; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; ON; w Eurasia [Introduced in North America; introduced also in Central America, South America, e Asia (Japan), South Africa, Australia]
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Discussion | Herbarium specimens of Cardamine hirsuta have been misidentified as C. oligosperma. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 469. | FNA vol. 7, p. 475. |
Parent taxa | Brassicaceae > tribe Cardamineae > Cardamine | Brassicaceae > tribe Cardamineae > Cardamine |
Sibling taxa | ||
Synonyms | Arabis bulbosa, Arabis rhomboidea, C. rhomboidea, C. rhomboidea var. hirsuta, C. rhomboidea var. parviflora, C. rhomboidea var. pilosa, Dentaria rhomboidea, Dracamine bulbosa | |
Name authority | (Schreber ex Muhlenberg) Britton: Prelim. Cat., 4. (1888) | Linnaeus: Sp. Pl. 2: 655. (1753) |
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