Cardamine blaisdellii |
Cardamine angulata |
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angle bitter-cress, angle-leaf bitter-cress, seaside bitter-cress |
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Habit | Perennials; usually glabrous throughout, rarely pilose. | Perennials; glabrous or sparsely pubescent. |
Rhizomes | cylindrical, slender, 0.7–1.5 mm diam. |
cylindrical, slender, to 2 mm diam. |
Stems | erect or ascending, unbranched, 0.5–2(–2.5) dm. |
erect, unbranched, (1.5–)2.5–8.5(–10) dm, sparsely to densely hirsute at base. |
Rhizomal leaves | pinnately 5- or 7-foliolate, 2.5–12 cm (not fleshy), leaflets petiolulate; petiole 1.5–4(–9) cm; lateral leaflets similar to terminal, or smaller and margins usually toothed, rarely entire; terminal leaflet (petiolule 0.15–0.3 cm) blade suborbicular to broadly obovate, 0.4–1.5 cm × 2.5–14 mm, base obtuse to subcordate, margins 3–5-toothed, (apiculate). |
3 (or 5)-foliolate, (4–)7–20(–22) cm, leaflets petiolulate or subsessile; petiole (2–)4–12(–14) cm; lateral leaflets subsessile, blade similar to terminal, larger or smaller in size; terminal leaflet petiolulate [(0.3–)0.5–1.5 cm], blade ovate to broadly lanceolate, 1.5–7(–9) cm, base usually cuneate, rarely subreniform or obtuse, margins 3–5(–7)-lobed or -toothed, surfaces puberulent. |
Cauline leaves | 1–3, (3–)5-foliolate (alternate), petiolate, leaflets petiolulate or subsessile; petiole 0.2–2(–6.5) cm, base not auriculate; lateral leaflets similar to terminal; terminal leaflet subsessile or petiolulate (to 0.2 cm), blade obovate to oblanceolate, 0.5–1.5 cm × 3–10 mm, base cuneate, margins 3-toothed or entire. |
(3 or) 4–8, 3 (or 5)-foliolate, petiolate, leaflets petiolulate or sessile; petiole 1–4 cm, base not auriculate; lateral leaflets sessile, blade similar to terminal, smaller, margins usually dentate, rarely entire; terminal leaflet sessile or petiolulate, blade broadly ovate to narrowly lanceolate, 2–7 cm × 6–40 mm, margins minutely pubescent. |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 2–3.5 × 2–2.5 mm, lateral pair slightly saccate basally; petals white, broadly obovate, 7–10 × 3–6 mm (clawed, apex rounded); filaments: median pairs 3–4 mm, lateral pair 2–3 mm; anthers oblong, 1–1.5 mm. |
sepals oblong, 2.5–4 × 1.3–2 mm, lateral pair saccate basally; petals usually white, rarely pinkish, obovate, 8–15 × 4–8 mm (clawed, apex rounded or emarginate); filaments: median pairs 3.5–6 mm, lateral pair 2–3.5 mm; anthers oblong, 0.8–1.2 mm. |
Fruiting pedicels | erect to ascending, 0.7–22 mm. |
ascending to divaricate, (9–)12–25 mm. |
Fruits | linear, 1.6–4 cm × 1–1.3 mm; ovules 14–24 per ovary; style 0.7–3 mm. |
linear, 1.5–3.2 cm × 1.4–2 mm; ovules 10–16 per ovary; style (0.5–)1–4 mm. |
Seeds | brown, oblong, ca. 1.5 × 1 mm. |
dark brown, oblong, 1.8–2.3 × 1–1.2 mm. |
2n | = 28, 42. |
= 40. |
Cardamine blaisdellii |
Cardamine angulata |
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Phenology | Flowering Jul–Aug. | Flowering Apr–Jun. |
Habitat | Moist streamsides, meadows, river gravel, mesic grounds, wet tundra, moist humus, scree slopes, calcareous fellfields | Moist ground, stream banks, swampy or damp woods, thickets, wet meadows |
Elevation | 50-1000 m (200-3300 ft) | 0-900 m (0-3000 ft) |
Distribution |
AK; n Europe; e Europe (Russian Far East, Siberia) |
AK; CA; OR; WA; BC
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Discussion | Cardamine blaisdellii was treated by R. C. Rollins (1993) as a synonym of C. microphylla, but the morphological differences (see key) and molecular data (R. B. Jørgensen et al., 2008) clearly demonstrate that they are distinct. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 468. | FNA vol. 7, p. 467. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. microphylla subsp. blaisdellii, C. microphylla var. blaisdellii | C. angulata var. alba, C. angulata var. hirsuta, C. angulata var. pentaphylla, Dentaria grandiflora |
Name authority | Eastwood: Bot. Gaz. 33: 146. (1902) | Hooker: Fl. Bor.-Amer. 1: 44. (1829) |
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