Cardamine bellidifolia |
Cardamine concatenata |
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alpine bitter-cress |
cut-leaf toothwort |
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Habit | Perennials (somewhat cespitose, subscapose); glabrous throughout. | Perennials; usually sparsely pubescent, sometimes glabrous. |
Rhizomes | absent (caudex usually with slender, rarely stout, rhizomelike branches). |
(tuberiform, fragile), moniliform, segments fusiform, 2–10(–20) mm diam. (fleshy). |
Stems | erect to ascending, branched (several), 0.1–0.8(–1.4) dm, glabrous. |
erect, unbranched, (1–)2–4(–5.5) dm, glabrous or pubescent distally. |
Basal leaves | rosulate, simple, (0.6–)1.2–5(–7) cm; petiole (3–)1–3.5(–5.5) cm; blade usually ovate to oblong, rarely oblanceolate to obovate, (0.4–)0.8–1.7(–2.5) cm × (2–)5–10(–16) mm, base cuneate to obtuse, margins usually entire, rarely repand or obtusely small-toothed (apex obtuse). |
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Rhizomal leaves | 3-foliolate, (7–)10–20(–30) cm, leaflets sessile; petiole (4–)7–18(–25) cm; lateral leaflets similar to terminal, blade sometimes smaller; terminal leaflet blade oblong, lanceolate, oblanceolate, or linear, 2.5–6 cm, base cuneate, margins coarsely dentate to incised, laciniate, or 3-lobed (lobes usually toothed to incised, rarely entire, surfaces puberulent or not, trichomes 0.2–0.3 mm). |
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Cauline leaves | 0 or 1 (or 2), simple, shortly petiolate; petiole base not auriculate; blade similar to basal leaves, much smaller. |
(2 or) 3, 3-foliolate (usually whorled or opposite, rarely alternate, similar in morphology to rhizomal leaves), petiolate, leaflets petiolulate or subsessile; petiole (1–)1.5–6(–7.5) cm, base not auriculate; lateral leaflets sessile, blade similar to terminal, sometimes smaller; terminal leaflet subsessile or petiolulate (to 3 cm), blade lanceolate, linear, or oblanceolate, (3–)4–10(–12) cm × (3–)5–20(–25) mm, margins usually coarsely dentate to incised, rarely subentire (margins minutely puberulent). |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, 2–3(–4) × 0.8–1.5(–2) mm, lateral pair not saccate basally; petals white, oblanceolate, 4–5.5(–7) × 1.3–2(–2.5) mm (not clawed, apex rounded or emarginate); filaments: median pairs 2.5–4 mm, lateral pair 2–3 mm; anthers oblong, 0.5–0.9 mm. |
sepals oblong, (4–)5–8 × 2–4 mm, lateral pair slightly saccate basally; petals white to pale pink, oblanceolate, (8–)10–20 × (3–)4–7(–9) mm, (short-clawed, apex rounded); filaments: median pairs 8–12 mm, lateral pair 6–8 mm; anthers oblong-linear, 1.5–2.5 mm. |
Fruiting pedicels | ascending to erect, 3–6(–8) mm. |
ascending to divaricate, (6–)10–27(–33) mm. |
Fruits | linear, (0.8–)1.3–2.8(–3.7) cm × 1.3–2 mm; ovules 8–18 per ovary; style 0.5–3 mm. |
linear-lanceolate, (2–)2.5–3.8(–4.8) cm × 1.5–3 mm; (valves glabrous or sparsely pubescent); ovules 10–14 per ovary; style (2–)5–9(–12) mm. |
Seeds | brown, oblong, 1.5–2 × 0.9–1.2 mm. |
brown, oblong, 1.6–3 × 1.8–2 mm. |
2n | = 16. |
= 128–256. |
Cardamine bellidifolia |
Cardamine concatenata |
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Phenology | Flowering Jun–Sep. | Flowering Feb–May. |
Habitat | Mossy areas, tundra, marshes at stream headwaters, cliffs, talus slopes, barren chert slopes, moist rock crevices, rocky slopes, alpine streams, sandy beaches, moist rocky streambeds | Wooded bottoms and bluffs, rich woods, limestone cliffs and outcrops, rocky banks, mesic forests, moist areas with leaf litter, floodplain woods |
Elevation | 0-2300 m (0-7500 ft) | 0-1000 m (0-3300 ft) |
Distribution |
AK; CA; ME; NH; OR; WA; AB; BC; LB; NT; NU; QC; YT; Greenland; n Europe; Asia (Russian Far East, Siberia)
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AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; VT; WI; WV; ON; QC
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Source | FNA vol. 7, p. 468. | FNA vol. 7, p. 470. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. bellidifolia var. beringensis, C. bellidifolia var. laxa, C. bellidifolia var. pachyphylla, C. bellidifolia var. pinnatifida | Dentaria concatenata, C. laciniata var. integra, C. laciniata var. lasiocarpa, Dentaria laciniata, Dentaria laciniata var. alterna, Dentaria laciniata var. coalescens, Dentaria laciniata var. integra, Dentaria laciniata var. latifolia, Dentaria laciniata var. opposita |
Name authority | Linnaeus: Sp. Pl. 2: 654. (1753) | (Michaux) O. Schwarz: Repert. Spec. Nov. Regni Veg. 46: 188. (1939) |
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