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pale yellow sun cup

California sun cup, southern suncup

Habit Herbs annual, appearing conspicuously grayish, densely strigillose, sometimes also glandular puberulent distally. Herbs annual, rarely short-lived perennial, usually villous, sometimes strigillose.
Stems

usually with decumbent lateral branches from basal rosette, 5–60 cm.

1–several from base, ascending or decumbent, to 80 cm.

Leaves

1–5(–11) × 0.2–0.7(–1.4) cm;

petiole 0–0.2(–0.4) cm, distal ones sessile;

blade lanceolate to narrowly ovate, base often cuneate to truncate, sometimes attenuate, margins sparsely denticulate, apex acute to obtuse.

1.2–12 × 0.2–1.5 cm;

petiole 0–4 cm, distal ones 0–0.3 cm;

blade (basal) narrowly elliptic or (cauline) usually narrowly lanceolate or lanceolate, rarely linear, base (basal) narrowly cuneate, (cauline) cuneate or subcordate, margins usually sparsely and inconspicuously denticulate, apex acute.

Flowers

opening near sunrise;

floral tube 1–4.2 mm;

sepals (1.5–)2.5–8 mm;

petals yellow, sometimes with 1–3 red dots basally, (2–)3.5–13 mm; episepalous filaments (0.5–)1.5–6.5 mm, epipetalous filaments (0.2–)0.5–3.8 mm, anthers (0.4–)0.8–2.2 mm, less than 5% of pollen grains 4- or 5-pored;

style (2.1–)3–10.5 mm, stigma surrounded by at least anthers of longer stamens, often by both sets, at anthesis.

opening near sunrise;

floral tube 2–5(–7.5) mm;

sepals (2.3–)5–8(–11) mm;

petals yellow, each usually with 1 bright red dot, rarely 2, near base, (4.2–)7–15 mm; episepalous filaments (1–)1.5–3.5 mm, epipetalous filaments (0.5–)1–2.5 mm, anthers (0.5–)1.3–2(–2.5) mm, less than 5% of pollen grains 4- or 5-pored;

style (5.5–)7–12 mm, stigma exserted beyond anthers at anthesis.

Capsules

usually 1–3-coiled spiral, subterete in living material, 4-angled when dry, 13–24 × 0.7–1.2 mm.

straight or somewhat contorted, weakly 4-angled, 12–40 × 1.5–2.5 mm.

Seeds

1–1.5 mm.

0.9–1 mm.

2n

= 14.

Camissoniopsis pallida

Camissoniopsis bistorta

Phenology Flowering Mar–Jun.
Habitat Sandy or clayey soils, coastal strands, grasslands, coastal sage scrub, chaparral, oak woodlands, margins of Sonoran and Mojave deserts, rarely higher elevation meadows.
Elevation 0–1600(–2600) m. (0–5200(–8500) ft.)
Distribution
w United States; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
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[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

P. H. Raven (1969) determined Camissoniopsis pallida to be self-compatible and primarily autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Camissoniopsis bistorta occurs in California from Ventura County south and east through the counties of southern Los Angeles, southwestern San Bernardino, Orange, western Riverside, and the western two-thirds of San Diego, reaching the margins of the desert in San Bernardino and San Diego counties, and southward in cismontane Baja California to Ojos Negros and San Vicente. The species occurs at exceptionally high elevations in the Santa Ana drainage of the San Bernardino Mountains. P. H. Raven (1969) indicated that there were occasional apparent hybrids between C. cheiranthifolia subsp. suffruticosa and C. bistorta occurring in intermediate habitats in areas where the two species co-occur. He determined that C. bistorta is self-incompatible.

Camissoniopsis bistorta was apparently introduced with stream gravel in 1959 in Goleta Marsh, Santa Barbara, California, and on ballast heaps at Nanaimo, Vancouver Island, British Columbia, in 1893. It has apparently not persisted at either site.

Oenothera heterophylla Nuttall ex Hooker & Arnott (1839), not Spach (1836), is an illegitimate name that pertains to Camissoniopsis bistorta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Floral tube 1–3 mm; petals (2–)3.5–6(–8) mm; styles (2.1–)3–6.5 mm.
subsp. pallida
1. Floral tube 3.8–4.2 mm; petals 6.5–13 mm; styles 6.5–10.5 mm.
subsp. hallii
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis
Sibling taxa
C. bistorta, C. cheiranthifolia, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. ignota, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. robusta
C. cheiranthifolia, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. ignota, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. pallida, C. robusta
Subordinate taxa
C. pallida subsp. hallii, C. pallida subsp. pallida
Synonyms Sphaerostigma pallidum, Camissonia pallida, Oenothera abramsii, O. micrantha var. abramsii Oenothera bistorta, Camissonia bistorta, O. bistorta var. veitchiana, Sphaerostigma bistortum, S. bistortum var. veitchianum, S. veitchianum
Name authority (Abrams) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 205. (2007) (Nuttall ex Torrey & A. Gray) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007)
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