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miniature suncup, Spencer primrose

Hardham's evening-primrose

Habit Herbs annual, densely villous, more densely so distally, also rarely glandular puberulent distally. Herbs annual, villous, also glandular puberulent distally.
Stems

arising from base, usually decumbent, rarely with 1 erect, 15–60 cm.

erect, with 1 or more branches from basal rosette, to 60 cm.

Leaves

1–12 × 0.2–1.7 cm;

petiole 0–2 cm, distal ones 0–0.5 cm;

blade (basal) narrowly elliptic, (cauline) narrowly elliptic-lanceolate to lanceolate, base (basal) narrowly cuneate, (cauline) rounded, margins denticulate, apex acute.

1–12 × 0.4–1.8 cm;

subsessile;

blade lanceolate, narrowly elliptic, or narrowly ovate, base truncate, margins dentate, apex acute.

Flowers

opening near sunrise;

floral tube 1.2–2 mm;

sepals 1–2.2(–2.5) mm;

petals yellow, sometimes with 1 or 2 red dots near base, 1.5–3.5(–4.5) mm; episepalous filaments 0.8–1.5 mm, epipetalous filaments 0.5–0.8(–1) mm, anthers 0.4–0.6 mm, less than 5% of pollen grains 4- or 5-pored;

style 2–3.5 mm, stigma surrounded by anthers at anthesis.

opening near sunrise;

floral tube 1.7–2 mm;

sepals 1.8–3.2 mm;

petals yellow, immaculate, 2–4 mm; episepalous filaments 1.5–2 mm, epipetalous filaments 1–1.5 mm, anthers 0.7 mm, 70–100% of pollen grains 4- or 5-pored;

style 3–4 mm, stigma surrounded by anthers at anthesis.

Capsules

straight or curved, equal to or slightly more than 1 complete spiral, subterete in living material, 4-angled when dry, 13–20(–25) × 1.1–1.2(–1.8) mm.

straight or 1-coiled, subterete in living material, obscurely 4-angled when dry, 13–25 × 1.3–1.6 mm.

Seeds

0.7–1.1 mm.

0.7–1.1 mm.

2n

= 14.

= 42.

Camissoniopsis micrantha

Camissoniopsis hardhamiae

Phenology Flowering (Jan–)Mar–Jun(–Sep). Flowering Mar–May.
Habitat Coastal strand, coastal sage scrub, chaparral. Sandy soils, limestone, disturbed oak woodlands.
Elevation 0–300(–800) m. (0–1000(–2600) ft.) 150–1000 m. (500–3300 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Camissoniopsis micrantha occurs from the vicinity of Bodega Bay, Sonoma County, near Lower Lake, Lake County, and near Rio Vista, Sacramento County, south in the Coast Ranges to the Los Angeles Basin and the northern edge of San Diego County; also on San Miguel, Santa Rosa, Santa Cruz, and Santa Catalina islands. The species was introduced, apparently on ballast heaps, at Nanaimo, Vancouver Island, British Columbia (Macoun s.n. in 1893, NMC). It has apparently not persisted in this area. P. H. Raven (1969) determined C. micrantha to be self-compatible and primarily autogamous. Excluded populations are now recognized as C. hirtella, C. ignota, C. lewisii, and C. pallida.

Oenothera hirta Link (1821), not Linnaeus (1759), is an illegitimate name that pertains to Camissoniopsis micrantha.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Camissoniopsis hardhamiae is narrowly endemic to the Outer South Coast Ranges. Populations are very local, known only from a few localities in sandy soil in disturbed oak woodland, southernmost Monterey to central San Luis Obispo County. P. H. Raven (1969) determined C. hardhamiae to be self-compatible and primarily autogamous. The species is apparently a hexaploid derived via hybridization between the tetraploid C. intermedia (2n = 28) and the diploid C. micrantha (2n = 14).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis
Sibling taxa
C. bistorta, C. cheiranthifolia, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. ignota, C. intermedia, C. lewisii, C. luciae, C. pallida, C. robusta
C. bistorta, C. cheiranthifolia, C. confusa, C. guadalupensis, C. hirtella, C. ignota, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. pallida, C. robusta
Synonyms Oenothera micrantha, Camissonia micrantha, Holostigma micranthum, S. micranthum Camissonia hardhamiae
Name authority (Hornemmann ex Sprengel) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 205. (2007) (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007)
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