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Lewis' evening-primrose

beach evening-primrose, beach suncup

Habit Herbs annual, villous, also glandular puberulent distally. Herbs short-lived perennial, sometimes woody at base, usually densely strigillose throughout, rarely glabrous, also villous distally.
Stems

usually several, decumbent, rarely 1 erect stem, 30–60 cm.

prostrate, decumbent, or ascending from base, to 60(–130) cm.

Leaves

1–8 × 0.2–1.1 cm;

petiole 0–3 cm;

blade narrowly lanceolate-elliptic, base cuneate or subcordate, margins denticulate, apex acute.

0.5–5 × 0.3–2.2 cm;

petiole 0–1.5(–2.5) cm, distal ones to 1 cm;

blade narrowly ovate, base attenuate, cuneate, or cordate, margins sparsely serrulate, apex acute.

Flowers

opening near sunrise;

floral tube 1.5–4 m;

sepals 1.7–3.4 mm;

petals yellow, with 1 or 2 red dots basally, 2.5–5.5 mm; episepalous filaments 2–2.8 mm, epipetalous filaments 1–1.7 mm, anthers 0.7–1.2 mm, less than 5% of pollen grains 4- or 5-pored;

style 2.8–4.5 mm, stigma surrounded by anthers at anthesis.

opening near sunrise;

floral tube 2.1–8.5 mm;

sepals 4–11.5 mm;

petals yellow, often red-dotted near base, 6–20 mm; episepalous filaments 2.8–8 mm, epipetalous filaments 1.5–6 mm, anthers 1–3 mm, less than 5% of pollen grains 4- or 5-pored;

style 6–23 mm, stigma surrounded by or exserted beyond anthers at anthesis.

Capsules

usually loosely 1-coiled, conspicuously 4-angled in living material, 13–20 × 1.8–2.2 mm.

often coiled in 1–2 spirals, 4-angled, 10–25 × 2–2.5 mm.

Seeds

0.7–0.8 mm.

1.2–1.3 mm.

2n

= 14.

Camissoniopsis lewisii

Camissoniopsis cheiranthifolia

Phenology Flowering Mar–May(–Sep).
Habitat Open sandy and clayey grasslands, coastal dunes and beaches.
Elevation 0–300 m. [0–1000 ft.]
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
w United States; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Camissoniopsis lewisii occurs from Point Dume and the Los Angeles Basin, Los Angeles County, south to Cardon Grande at the northern edge of Baja California Sur. P. H. Raven (1969) determined C. lewisii to be self-compatible and primarily autogamous, and suggested that this coastal Camissoniopsis may have been derived more or less directly from coastal populations of C. bistorta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Camissoniopsis cheiranthifolia occurs on slopes and dunes along the immediate coast and on islands from Coos Bay, Curry County, Oregon, to the vicinity of San Quintín, Baja California; it is also known from the east shore of San Francisco Bay and locally on sand dunes along the lower Sacramento River, California, 0–100 m. P. H. Raven (1969) determined C. cheiranthifolia to be self-incompatible (some populations in subsp. suffruticosa) or self-compatible (both subspecies) and apparently pollinated by oligolectic bees of Andrena subg. Onagrandrena (Raven); Raven subdivided the species into two intergrading subspecies.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants primarily herbaceous; petals 6–11 mm; style 6–9 mm; stigma usually surrounded by anthers.
subsp. cheiranthifolia
1. Plants woody at base; petals (10–)12–20 mm; style 13–23 mm; stigma exserted beyond anthers at anthesis.
subsp. suffruticosa
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis
Sibling taxa
C. bistorta, C. cheiranthifolia, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. ignota, C. intermedia, C. luciae, C. micrantha, C. pallida, C. robusta
C. bistorta, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. ignota, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. pallida, C. robusta
Subordinate taxa
C. cheiranthifolia subsp. cheiranthifolia, C. cheiranthifolia subsp. suffruticosa
Synonyms Camissonia lewisii Oenothera cheiranthifolia, Agassizia cheiranthifolia, Camissonia cheiranthifolia, Holostigma cheiranthifolium, Sphaerostigma cheiranthifolium
Name authority (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 205. (2007) (Hornemann ex Sprengel) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007)
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