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hairy sun cup, Santa Cruz Island suncup

beach evening-primrose, beach suncup

Habit Herbs annual, densely villous throughout, also glandular puberulent distally. Herbs short-lived perennial, sometimes woody at base, usually densely strigillose throughout, rarely glabrous, also villous distally.
Stems

erect, with 1 or more ascending branches from near base, to 60 cm.

prostrate, decumbent, or ascending from base, to 60(–130) cm.

Leaves

1–11 × 0.3–2.1 cm;

petiole 0–5 cm, distal ones 0–0.5 cm;

blade lanceolate to ovate, sometimes elliptic-ovate or ovate distally, base cordate to truncate, sometimes cuneate or attenuate, margins dentate, apex acute.

0.5–5 × 0.3–2.2 cm;

petiole 0–1.5(–2.5) cm, distal ones to 1 cm;

blade narrowly ovate, base attenuate, cuneate, or cordate, margins sparsely serrulate, apex acute.

Flowers

opening near sunrise;

floral tube 1–3 mm;

sepals 2.5–6 mm;

petals yellow, sometimes red-dotted near base, 2–9 mm, sometimes with a tooth arising from emarginate apex; episepalous filaments 1.2–6 mm, epipetalous filaments 0.5–3 mm, anthers 0.4–1 mm, less than 5% of pollen grains 4- or 5-pored;

style 2–8 mm, stigma surrounded by anthers at anthesis.

opening near sunrise;

floral tube 2.1–8.5 mm;

sepals 4–11.5 mm;

petals yellow, often red-dotted near base, 6–20 mm; episepalous filaments 2.8–8 mm, epipetalous filaments 1.5–6 mm, anthers 1–3 mm, less than 5% of pollen grains 4- or 5-pored;

style 6–23 mm, stigma surrounded by or exserted beyond anthers at anthesis.

Capsules

1–2-coiled spiral, subterete in living material, 4-angled when dry, 13–20(–25) × 0.7–0.9 mm.

often coiled in 1–2 spirals, 4-angled, 10–25 × 2–2.5 mm.

Seeds

1–1.2 mm.

1.2–1.3 mm.

2n

= 14.

Camissoniopsis hirtella

Camissoniopsis cheiranthifolia

Phenology Flowering (Jan–)Mar–Jul(–Nov).
Habitat Brushy hills and slopes, on burns.
Elevation 0–2300 m. (0–7500 ft.)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
w United States; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Camissoniopsis hirtella occurs from Amador and Trinity counties southward in the Coast Ranges and Sierra Nevada of California to the Sierra de Juárez and Sierra San Pedro Mártir, Baja California, usually away from the immediate coast and barely reaching the margins of the desert. P. H. Raven (1969) determined C. hirtella to be self-compatible and primarily autogamous. The species occasionally hybridizes with C. ignota (Raven).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Camissoniopsis cheiranthifolia occurs on slopes and dunes along the immediate coast and on islands from Coos Bay, Curry County, Oregon, to the vicinity of San Quintín, Baja California; it is also known from the east shore of San Francisco Bay and locally on sand dunes along the lower Sacramento River, California, 0–100 m. P. H. Raven (1969) determined C. cheiranthifolia to be self-incompatible (some populations in subsp. suffruticosa) or self-compatible (both subspecies) and apparently pollinated by oligolectic bees of Andrena subg. Onagrandrena (Raven); Raven subdivided the species into two intergrading subspecies.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants primarily herbaceous; petals 6–11 mm; style 6–9 mm; stigma usually surrounded by anthers.
subsp. cheiranthifolia
1. Plants woody at base; petals (10–)12–20 mm; style 13–23 mm; stigma exserted beyond anthers at anthesis.
subsp. suffruticosa
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis
Sibling taxa
C. bistorta, C. cheiranthifolia, C. confusa, C. guadalupensis, C. hardhamiae, C. ignota, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. pallida, C. robusta
C. bistorta, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. ignota, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. pallida, C. robusta
Subordinate taxa
C. cheiranthifolia subsp. cheiranthifolia, C. cheiranthifolia subsp. suffruticosa
Synonyms Oenothera hirtella, Camissonia hirtella, O. hirta var. jonesii, O. micrantha var. hirtella, O. micrantha var. jonesii, O. micrantha var. reedii, Sphaerostigma arenicola, S. bistortum var. reedii, S. hirtellum, S. hirtellum var. montanum, S. micranthum var. jonesii Oenothera cheiranthifolia, Agassizia cheiranthifolia, Camissonia cheiranthifolia, Holostigma cheiranthifolium, Sphaerostigma cheiranthifolium
Name authority (Greene) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007) (Hornemann ex Sprengel) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007)
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