Camissoniopsis hardhamiae
Onagraceae tribe Onagreae
Hardham's evening-primrose
erect, with 1 or more branches from basal rosette, to 60 cm.
1–12 × 0.4–1.8 cm;
subsessile;
blade lanceolate, narrowly elliptic, or narrowly ovate, base truncate, margins dentate, apex acute.
alternate or basal;
stipules absent.
opening near sunrise;
floral tube 1.7–2 mm;
sepals 1.8–3.2 mm;
petals yellow, immaculate, 2–4 mm; episepalous filaments 1.5–2 mm, epipetalous filaments 1–1.5 mm, anthers 0.7 mm, 70–100% of pollen grains 4- or 5-pored;
style 3–4 mm, stigma surrounded by anthers at anthesis.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
straight or 1-coiled, subterete in living material, obscurely 4-angled when dry, 13–25 × 1.3–1.6 mm.
0.7–1.1 mm.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
= 42.
Camissoniopsis hardhamiae
Onagraceae tribe Onagreae
Camissoniopsis hardhamiae is narrowly endemic to the Outer South Coast Ranges. Populations are very local, known only from a few localities in sandy soil in disturbed oak woodland, southernmost Monterey to central San Luis Obispo County. P. H. Raven (1969) determined C. hardhamiae to be self-compatible and primarily autogamous. The species is apparently a hexaploid derived via hybridization between the tetraploid C. intermedia (2n = 28) and the diploid C. micrantha (2n = 14).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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