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beach evening-primrose, beach suncup

robust subcup, robust suncup

Habit Herbs short-lived perennial, sometimes woody at base, usually densely strigillose throughout, rarely glabrous, also villous distally. Herbs annual, villous, usually also glandular puberulent distally.
Stems

prostrate, decumbent, or ascending from base, to 60(–130) cm.

erect, with 1 or more ascending branches from base, to 60 cm.

Leaves

0.5–5 × 0.3–2.2 cm;

petiole 0–1.5(–2.5) cm, distal ones to 1 cm;

blade narrowly ovate, base attenuate, cuneate, or cordate, margins sparsely serrulate, apex acute.

1–8 × 1.5–2 cm;

subsessile;

blade narrowly lanceolate-elliptic, base cuneate to truncate, basal often attenuate, margins denticulate, apex acute.

Flowers

opening near sunrise;

floral tube 2.1–8.5 mm;

sepals 4–11.5 mm;

petals yellow, often red-dotted near base, 6–20 mm; episepalous filaments 2.8–8 mm, epipetalous filaments 1.5–6 mm, anthers 1–3 mm, less than 5% of pollen grains 4- or 5-pored;

style 6–23 mm, stigma surrounded by or exserted beyond anthers at anthesis.

opening near sunrise;

floral tube 1.8–3.7 mm;

sepals 2.6–4.2 mm;

petals yellow, usually with 1 or 2 red dots basally, 3.2–7 mm; episepalous filaments 1.8–3 mm, epipetalous filaments 1–1.5 mm, anthers 0.8–3 mm, 70–100% of pollen grains 4-pored;

style 3–6.2 mm, stigma surrounded by anthers at anthesis.

Capsules

often coiled in 1–2 spirals, 4-angled, 10–25 × 2–2.5 mm.

usually 1-coiled spiral, 4-angled, 14–25 × 1.5–2 mm.

Seeds

1.2–1.3 mm.

0.9–1.2 mm.

2n

= 42.

Camissoniopsis cheiranthifolia

Camissoniopsis robusta

Phenology Flowering (Jan–)Mar–Jun(–Sep).
Habitat Coastal sage, chaparral, disturbed or open places.
Elevation 0–600(–800) m. (0–2000(–2600) ft.)
Distribution
w United States; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Camissoniopsis cheiranthifolia occurs on slopes and dunes along the immediate coast and on islands from Coos Bay, Curry County, Oregon, to the vicinity of San Quintín, Baja California; it is also known from the east shore of San Francisco Bay and locally on sand dunes along the lower Sacramento River, California, 0–100 m. P. H. Raven (1969) determined C. cheiranthifolia to be self-incompatible (some populations in subsp. suffruticosa) or self-compatible (both subspecies) and apparently pollinated by oligolectic bees of Andrena subg. Onagrandrena (Raven); Raven subdivided the species into two intergrading subspecies.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Camissoniopsis robusta occurs in coastal San Diego County, California, and coastal northwestern Baja California, south to the vicinity of El Rosario; also on Guadalupe, San Clemente, Santa Catalina, and Santa Cruz (rare) islands. P. H. Raven (1969) determined C. robusta to be self-compatible and primarily autogamous. Based on the intermediate morphology of this hexaploid, Raven suggested that it was derived from two species with which it occurs nearly throughout its rather limited range, the tetraploid C. intermedia (2n = 28) and the diploid C. lewisii (2n = 14).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants primarily herbaceous; petals 6–11 mm; style 6–9 mm; stigma usually surrounded by anthers.
subsp. cheiranthifolia
1. Plants woody at base; petals (10–)12–20 mm; style 13–23 mm; stigma exserted beyond anthers at anthesis.
subsp. suffruticosa
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis
Sibling taxa
C. bistorta, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. ignota, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. pallida, C. robusta
C. bistorta, C. cheiranthifolia, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. ignota, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. pallida
Subordinate taxa
C. cheiranthifolia subsp. cheiranthifolia, C. cheiranthifolia subsp. suffruticosa
Synonyms Oenothera cheiranthifolia, Agassizia cheiranthifolia, Camissonia cheiranthifolia, Holostigma cheiranthifolium, Sphaerostigma cheiranthifolium Camissonia robusta
Name authority (Hornemann ex Sprengel) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007) (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 205. (2007)
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