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California sun cup, southern suncup

jurupa hills sun cup

Habit Herbs annual, rarely short-lived perennial, usually villous, sometimes strigillose. Herbs annual, strigillose, usually also sparsely villous, often also glandular puberulent distally.
Stems

1–several from base, ascending or decumbent, to 80 cm.

arising from base, usually decumbent, rarely with only 1, erect stem, 10–55 cm.

Leaves

1.2–12 × 0.2–1.5 cm;

petiole 0–4 cm, distal ones 0–0.3 cm;

blade (basal) narrowly elliptic or (cauline) usually narrowly lanceolate or lanceolate, rarely linear, base (basal) narrowly cuneate, (cauline) cuneate or subcordate, margins usually sparsely and inconspicuously denticulate, apex acute.

1.5–7 × 0.3–1.3 cm;

petiole (0–)0.2–2.5 cm, petiolate distally;

blade narrowly lanceolate, lanceolate, or narrowly elliptic, base attenuate, margins serrulate, apex acute.

Flowers

opening near sunrise;

floral tube 2–5(–7.5) mm;

sepals (2.3–)5–8(–11) mm;

petals yellow, each usually with 1 bright red dot, rarely 2, near base, (4.2–)7–15 mm; episepalous filaments (1–)1.5–3.5 mm, epipetalous filaments (0.5–)1–2.5 mm, anthers (0.5–)1.3–2(–2.5) mm, less than 5% of pollen grains 4- or 5-pored;

style (5.5–)7–12 mm, stigma exserted beyond anthers at anthesis.

opening near sunrise;

floral tube (1.1–)1.8–3 mm;

sepals 2.6–5.5 mm;

petals yellow, sometimes red-dotted near base, (3–)4–8 mm; episepalous filaments (1.2–)2.5–3.6 mm, epipetalous filaments (1–)1.3–2 mm, anthers (0.6–)0.8–1.6 mm, less than 5% of pollen grains 4- or 5-pored;

style (3–)4.5–7 mm, stigma surrounded by anthers at anthesis.

Capsules

straight or somewhat contorted, weakly 4-angled, 12–40 × 1.5–2.5 mm.

very slender, usually much contorted, irregularly to 5-coiled, rarely simply flexuous, terete in living material, 4-angled when dry, 20–30 × 0.8–1 mm.

Seeds

0.9–1 mm.

1.2–1.3 mm.

2n

= 14.

= 14.

Camissoniopsis bistorta

Camissoniopsis ignota

Phenology Flowering Mar–Jun. Flowering (Jan–)Mar–Apr(–Aug).
Habitat Sandy or clayey soils, coastal strands, grasslands, coastal sage scrub, chaparral, oak woodlands, margins of Sonoran and Mojave deserts, rarely higher elevation meadows. Clay or sandy soils, flats and slopes in coastal sage scrub or chap­arral, sandy soils in mountains.
Elevation 0–1600(–2600) m. [0–5200(–8500) ft.] 100–1100(–1500) m. [300–3600(–4900) ft.]
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Camissoniopsis bistorta occurs in California from Ventura County south and east through the counties of southern Los Angeles, southwestern San Bernardino, Orange, western Riverside, and the western two-thirds of San Diego, reaching the margins of the desert in San Bernardino and San Diego counties, and southward in cismontane Baja California to Ojos Negros and San Vicente. The species occurs at exceptionally high elevations in the Santa Ana drainage of the San Bernardino Mountains. P. H. Raven (1969) indicated that there were occasional apparent hybrids between C. cheiranthifolia subsp. suffruticosa and C. bistorta occurring in intermediate habitats in areas where the two species co-occur. He determined that C. bistorta is self-incompatible.

Camissoniopsis bistorta was apparently introduced with stream gravel in 1959 in Goleta Marsh, Santa Barbara, California, and on ballast heaps at Nanaimo, Vancouver Island, British Columbia, in 1893. It has apparently not persisted at either site.

Oenothera heterophylla Nuttall ex Hooker & Arnott (1839), not Spach (1836), is an illegitimate name that pertains to Camissoniopsis bistorta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Camissoniopsis ignota is most common in clay fields and slopes at low elevations, but occasional on sandy soil and higher in the mountains in the Coast Ranges and bordering valleys from Yolo County, California, south to the southern end of the Sierra San Miguel, in Baja California, usually away from the immediate coast and barely reaching the margins of the desert. P. H. Raven (1969) determined C. ignota to be self-compatible and primarily autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis Onagraceae > subfam. Onagroideae > tribe Onagreae > Camissoniopsis
Sibling taxa
C. cheiranthifolia, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. ignota, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. pallida, C. robusta
C. bistorta, C. cheiranthifolia, C. confusa, C. guadalupensis, C. hardhamiae, C. hirtella, C. intermedia, C. lewisii, C. luciae, C. micrantha, C. pallida, C. robusta
Synonyms Oenothera bistorta, Camissonia bistorta, O. bistorta var. veitchiana, Sphaerostigma bistortum, S. bistortum var. veitchianum, S. veitchianum Oenothera micrantha var. ignota, Camissonia ignota, O. hirta var. ignota, O. ignota
Name authority (Nuttall ex Torrey & A. Gray) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007) (Jepson) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 205. (2007)
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