Camissoniopsis bistorta |
Camissoniopsis hirtella |
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California sun cup, southern suncup |
hairy sun cup, Santa Cruz Island suncup |
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Habit | Herbs annual, rarely short-lived perennial, usually villous, sometimes strigillose. | Herbs annual, densely villous throughout, also glandular puberulent distally. |
Stems | 1–several from base, ascending or decumbent, to 80 cm. |
erect, with 1 or more ascending branches from near base, to 60 cm. |
Leaves | 1.2–12 × 0.2–1.5 cm; petiole 0–4 cm, distal ones 0–0.3 cm; blade (basal) narrowly elliptic or (cauline) usually narrowly lanceolate or lanceolate, rarely linear, base (basal) narrowly cuneate, (cauline) cuneate or subcordate, margins usually sparsely and inconspicuously denticulate, apex acute. |
1–11 × 0.3–2.1 cm; petiole 0–5 cm, distal ones 0–0.5 cm; blade lanceolate to ovate, sometimes elliptic-ovate or ovate distally, base cordate to truncate, sometimes cuneate or attenuate, margins dentate, apex acute. |
Flowers | opening near sunrise; floral tube 2–5(–7.5) mm; sepals (2.3–)5–8(–11) mm; petals yellow, each usually with 1 bright red dot, rarely 2, near base, (4.2–)7–15 mm; episepalous filaments (1–)1.5–3.5 mm, epipetalous filaments (0.5–)1–2.5 mm, anthers (0.5–)1.3–2(–2.5) mm, less than 5% of pollen grains 4- or 5-pored; style (5.5–)7–12 mm, stigma exserted beyond anthers at anthesis. |
opening near sunrise; floral tube 1–3 mm; sepals 2.5–6 mm; petals yellow, sometimes red-dotted near base, 2–9 mm, sometimes with a tooth arising from emarginate apex; episepalous filaments 1.2–6 mm, epipetalous filaments 0.5–3 mm, anthers 0.4–1 mm, less than 5% of pollen grains 4- or 5-pored; style 2–8 mm, stigma surrounded by anthers at anthesis. |
Capsules | straight or somewhat contorted, weakly 4-angled, 12–40 × 1.5–2.5 mm. |
1–2-coiled spiral, subterete in living material, 4-angled when dry, 13–20(–25) × 0.7–0.9 mm. |
Seeds | 0.9–1 mm. |
1–1.2 mm. |
2n | = 14. |
= 14. |
Camissoniopsis bistorta |
Camissoniopsis hirtella |
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Phenology | Flowering Mar–Jun. | Flowering (Jan–)Mar–Jul(–Nov). |
Habitat | Sandy or clayey soils, coastal strands, grasslands, coastal sage scrub, chaparral, oak woodlands, margins of Sonoran and Mojave deserts, rarely higher elevation meadows. | Brushy hills and slopes, on burns. |
Elevation | 0–1600(–2600) m. [0–5200(–8500) ft.] | 0–2300 m. [0–7500 ft.] |
Distribution |
CA; Mexico (Baja California)
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CA; Mexico (Baja California)
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Discussion | Camissoniopsis bistorta occurs in California from Ventura County south and east through the counties of southern Los Angeles, southwestern San Bernardino, Orange, western Riverside, and the western two-thirds of San Diego, reaching the margins of the desert in San Bernardino and San Diego counties, and southward in cismontane Baja California to Ojos Negros and San Vicente. The species occurs at exceptionally high elevations in the Santa Ana drainage of the San Bernardino Mountains. P. H. Raven (1969) indicated that there were occasional apparent hybrids between C. cheiranthifolia subsp. suffruticosa and C. bistorta occurring in intermediate habitats in areas where the two species co-occur. He determined that C. bistorta is self-incompatible. Camissoniopsis bistorta was apparently introduced with stream gravel in 1959 in Goleta Marsh, Santa Barbara, California, and on ballast heaps at Nanaimo, Vancouver Island, British Columbia, in 1893. It has apparently not persisted at either site. Oenothera heterophylla Nuttall ex Hooker & Arnott (1839), not Spach (1836), is an illegitimate name that pertains to Camissoniopsis bistorta. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Camissoniopsis hirtella occurs from Amador and Trinity counties southward in the Coast Ranges and Sierra Nevada of California to the Sierra de Juárez and Sierra San Pedro Mártir, Baja California, usually away from the immediate coast and barely reaching the margins of the desert. P. H. Raven (1969) determined C. hirtella to be self-compatible and primarily autogamous. The species occasionally hybridizes with C. ignota (Raven). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Oenothera bistorta, Camissonia bistorta, O. bistorta var. veitchiana, Sphaerostigma bistortum, S. bistortum var. veitchianum, S. veitchianum | Oenothera hirtella, Camissonia hirtella, O. hirta var. jonesii, O. micrantha var. hirtella, O. micrantha var. jonesii, O. micrantha var. reedii, Sphaerostigma arenicola, S. bistortum var. reedii, S. hirtellum, S. hirtellum var. montanum, S. micranthum var. jonesii |
Name authority | (Nuttall ex Torrey & A. Gray) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007) | (Greene) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007) |
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