Camissoniopsis bistorta |
Camissoniopsis hardhamiae |
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California sun cup, southern suncup |
Hardham's evening-primrose |
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Habit | Herbs annual, rarely short-lived perennial, usually villous, sometimes strigillose. | Herbs annual, villous, also glandular puberulent distally. |
Stems | 1–several from base, ascending or decumbent, to 80 cm. |
erect, with 1 or more branches from basal rosette, to 60 cm. |
Leaves | 1.2–12 × 0.2–1.5 cm; petiole 0–4 cm, distal ones 0–0.3 cm; blade (basal) narrowly elliptic or (cauline) usually narrowly lanceolate or lanceolate, rarely linear, base (basal) narrowly cuneate, (cauline) cuneate or subcordate, margins usually sparsely and inconspicuously denticulate, apex acute. |
1–12 × 0.4–1.8 cm; subsessile; blade lanceolate, narrowly elliptic, or narrowly ovate, base truncate, margins dentate, apex acute. |
Flowers | opening near sunrise; floral tube 2–5(–7.5) mm; sepals (2.3–)5–8(–11) mm; petals yellow, each usually with 1 bright red dot, rarely 2, near base, (4.2–)7–15 mm; episepalous filaments (1–)1.5–3.5 mm, epipetalous filaments (0.5–)1–2.5 mm, anthers (0.5–)1.3–2(–2.5) mm, less than 5% of pollen grains 4- or 5-pored; style (5.5–)7–12 mm, stigma exserted beyond anthers at anthesis. |
opening near sunrise; floral tube 1.7–2 mm; sepals 1.8–3.2 mm; petals yellow, immaculate, 2–4 mm; episepalous filaments 1.5–2 mm, epipetalous filaments 1–1.5 mm, anthers 0.7 mm, 70–100% of pollen grains 4- or 5-pored; style 3–4 mm, stigma surrounded by anthers at anthesis. |
Capsules | straight or somewhat contorted, weakly 4-angled, 12–40 × 1.5–2.5 mm. |
straight or 1-coiled, subterete in living material, obscurely 4-angled when dry, 13–25 × 1.3–1.6 mm. |
Seeds | 0.9–1 mm. |
0.7–1.1 mm. |
2n | = 14. |
= 42. |
Camissoniopsis bistorta |
Camissoniopsis hardhamiae |
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Phenology | Flowering Mar–Jun. | Flowering Mar–May. |
Habitat | Sandy or clayey soils, coastal strands, grasslands, coastal sage scrub, chaparral, oak woodlands, margins of Sonoran and Mojave deserts, rarely higher elevation meadows. | Sandy soils, limestone, disturbed oak woodlands. |
Elevation | 0–1600(–2600) m. [0–5200(–8500) ft.] | 150–1000 m. [500–3300 ft.] |
Distribution |
CA; Mexico (Baja California)
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CA |
Discussion | Camissoniopsis bistorta occurs in California from Ventura County south and east through the counties of southern Los Angeles, southwestern San Bernardino, Orange, western Riverside, and the western two-thirds of San Diego, reaching the margins of the desert in San Bernardino and San Diego counties, and southward in cismontane Baja California to Ojos Negros and San Vicente. The species occurs at exceptionally high elevations in the Santa Ana drainage of the San Bernardino Mountains. P. H. Raven (1969) indicated that there were occasional apparent hybrids between C. cheiranthifolia subsp. suffruticosa and C. bistorta occurring in intermediate habitats in areas where the two species co-occur. He determined that C. bistorta is self-incompatible. Camissoniopsis bistorta was apparently introduced with stream gravel in 1959 in Goleta Marsh, Santa Barbara, California, and on ballast heaps at Nanaimo, Vancouver Island, British Columbia, in 1893. It has apparently not persisted at either site. Oenothera heterophylla Nuttall ex Hooker & Arnott (1839), not Spach (1836), is an illegitimate name that pertains to Camissoniopsis bistorta. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Camissoniopsis hardhamiae is narrowly endemic to the Outer South Coast Ranges. Populations are very local, known only from a few localities in sandy soil in disturbed oak woodland, southernmost Monterey to central San Luis Obispo County. P. H. Raven (1969) determined C. hardhamiae to be self-compatible and primarily autogamous. The species is apparently a hexaploid derived via hybridization between the tetraploid C. intermedia (2n = 28) and the diploid C. micrantha (2n = 14). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Oenothera bistorta, Camissonia bistorta, O. bistorta var. veitchiana, Sphaerostigma bistortum, S. bistortum var. veitchianum, S. veitchianum | Camissonia hardhamiae |
Name authority | (Nuttall ex Torrey & A. Gray) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007) | (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007) |
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