Camissoniopsis bistorta |
Camissoniopsis confusa |
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California sun cup, southern suncup |
San Bernardino sun cup |
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Habit | Herbs annual, rarely short-lived perennial, usually villous, sometimes strigillose. | Herbs annual, densely villous, often also strigillose, at least sparsely villous and glandular puberulent on stems distally and on inflorescences. |
Stems | 1–several from base, ascending or decumbent, to 80 cm. |
erect, with multiple branches, rarely with 1 stem, to 70 cm. |
Leaves | 1.2–12 × 0.2–1.5 cm; petiole 0–4 cm, distal ones 0–0.3 cm; blade (basal) narrowly elliptic or (cauline) usually narrowly lanceolate or lanceolate, rarely linear, base (basal) narrowly cuneate, (cauline) cuneate or subcordate, margins usually sparsely and inconspicuously denticulate, apex acute. |
1–6 × 0.4–2 cm; petiole 0–3 cm, distal ones sessile; blade lanceolate or narrowly ovate, base round or truncate, margins sparsely denticulate, apex long-acuminate. |
Flowers | opening near sunrise; floral tube 2–5(–7.5) mm; sepals (2.3–)5–8(–11) mm; petals yellow, each usually with 1 bright red dot, rarely 2, near base, (4.2–)7–15 mm; episepalous filaments (1–)1.5–3.5 mm, epipetalous filaments (0.5–)1–2.5 mm, anthers (0.5–)1.3–2(–2.5) mm, less than 5% of pollen grains 4- or 5-pored; style (5.5–)7–12 mm, stigma exserted beyond anthers at anthesis. |
opening near sunrise; floral tube (1.8–)2–3.8 mm; sepals (1.5–)3.2–8.5 mm; petals yellow, usually with 1 or 2 red dots basally, (2.5–)5–10.5 mm; episepalous filaments (1.2–)2.5–4.5 mm, epipetalous filaments (0.8–)1.5–2.5 mm, anthers (0.4–)0.8–1.5 mm, less than 5% of pollen grains 4- or 5-pored; style (2.5–)4.5–7.5 mm, stigma surrounded by anthers at anthesis. |
Capsules | straight or somewhat contorted, weakly 4-angled, 12–40 × 1.5–2.5 mm. |
straight or 1–2-coiled spiral, subterete in living material, 4-angled when dry, 13–23 × 0.9–1.2 mm. |
Seeds | 0.9–1 mm. |
0.7–1.1 mm. |
2n | = 14. |
= 14. |
Camissoniopsis bistorta |
Camissoniopsis confusa |
|
Phenology | Flowering Mar–Jun. | Flowering Mar–Jun(–Jul). |
Habitat | Sandy or clayey soils, coastal strands, grasslands, coastal sage scrub, chaparral, oak woodlands, margins of Sonoran and Mojave deserts, rarely higher elevation meadows. | Dry inland slopes, chaparral. |
Elevation | 0–1600(–2600) m. [0–5200(–8500) ft.] | 300–2000 m. [1000–6600 ft.] |
Distribution |
CA; Mexico (Baja California)
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AZ; CA
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Discussion | Camissoniopsis bistorta occurs in California from Ventura County south and east through the counties of southern Los Angeles, southwestern San Bernardino, Orange, western Riverside, and the western two-thirds of San Diego, reaching the margins of the desert in San Bernardino and San Diego counties, and southward in cismontane Baja California to Ojos Negros and San Vicente. The species occurs at exceptionally high elevations in the Santa Ana drainage of the San Bernardino Mountains. P. H. Raven (1969) indicated that there were occasional apparent hybrids between C. cheiranthifolia subsp. suffruticosa and C. bistorta occurring in intermediate habitats in areas where the two species co-occur. He determined that C. bistorta is self-incompatible. Camissoniopsis bistorta was apparently introduced with stream gravel in 1959 in Goleta Marsh, Santa Barbara, California, and on ballast heaps at Nanaimo, Vancouver Island, British Columbia, in 1893. It has apparently not persisted at either site. Oenothera heterophylla Nuttall ex Hooker & Arnott (1839), not Spach (1836), is an illegitimate name that pertains to Camissoniopsis bistorta. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Camissoniopsis confusa occurs in California from the La Panza Range of central San Luis Obispo County south through the Coast Ranges to the San Bernardino Mountains and southern San Diego County; also in central Arizona (westernmost Gila, Maricopa, and northern Pinal counties). P. H. Raven (1969) determined C. confusa to be self-compatible and primarily autogamous. The species apparently is a tetraploid derived via hybridization between two diploid (2n = 14) species, C. hirtella and C. pallida. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Oenothera bistorta, Camissonia bistorta, O. bistorta var. veitchiana, Sphaerostigma bistortum, S. bistortum var. veitchianum, S. veitchianum | Camissonia confusa |
Name authority | (Nuttall ex Torrey & A. Gray) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007) | (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 204. (2007) |
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