Calystegia catesbeiana
Calystegia
Catesby's false bindweed
bindweed, false bindweed, morning-glory
pubescent to tomentose, hairs usually whitish.
usually twining-climbing, sometimes proximally erect, distally twining-climbing, to 40–200(–300) cm.
usually twining-climbing, sometimes ascending, decumbent, erect, procumbent, prostrate, or trailing, rarely almost absent, usually hairy, hairs not branched, glandular, or stellate, sometimes glabrate, glabrescent, or glabrous.
blade elliptic-ovate, to 120 × 50 mm, base lobed, lobes obtuse or rounded, to 20 mm.
petiolate;
blade usually cordate, elliptic, linear, oblong, oblong-hastate, orbiculate, oval, ovate, reniform, triangular, or triangular-hastate, rarely palmately 7–9-lobed, (15–)20–130 mm, base usually lobed or truncate, sometimes cuneate, surfaces glabrate, glabrescent, glabrous, ± pilose, tomentose, tomentulose, or villous.
immediately subtending sepals, lanceolate, 12–34 × 10–22 mm, proximally ± keeled, margins ± enfolding sepals, apex acute.
usually axillary, rarely terminal, compound cymes, bracteate;
peduncles 1(–4)-flowered.
sepals 11–17 mm;
corolla white, 44–64(–70) mm.
sepals ± elliptic, lanceolate, lance-ovate, oblong, oblong-ovate, oval, or ovate, (5–)8–15(–25) mm;
corolla usually white, sometimes cream, pink, purple, red, or yellow, campanulate to funnelform, (20–)25–60(–73)[–88] mm, limb entire or 5-lobed or -angled, rarely multilobed;
ovary 1-locular, sometimes with partial septum;
styles 1;
stigma lobes 2, linear to oblong, apices blunt.
capsular, ± globose, dehiscence irregular.
(1–)2–4, pyramidal to subglobose or trigonous, glabrous, papillate, smooth, or reticulate.
= 12.
Calystegia catesbeiana
Calystegia
Subspecies 2 (2 in the flora).
Plants of Calystegia catesbeiana, especially subsp. catesbeiana, often have been misidentified as C. sepium because of their climbing habit.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species ca. 30 (20 in the flora).
H. Hallier (1893) and W. H. Lewis and R. L. Oliver (1965) summarized arguments for treating Calystegia and Convolvulus as distinct genera. In a molecular analysis, M. A. Carine et al. (2004) found Calystegia nested within Convolvulus.
Species delimitation is problematic throughout Calystegia, with geographic and morphological intergradation between taxa, and often arbitrary limits have to be adopted to avoid impractically broad species. Hybridization is common where species overlap geographically. It is difficult to pinpoint any species which is not taxonomically subdivided and which does not intergrade or hybridize with others.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Stems usually weakly twining-climbing, sometimes proximally erect, to 40(–100) cm; leaf blades to 60 mm, basal lobes obtuse or rounded, to 11 mm, surfaces usually pubescent, abaxial rarely whitish. | subsp. catesbeiana |
1. Stems twining-climbing to 200(–300) cm; leaf blades to 120 mm, basal lobes rounded, 9–20 mm, surfaces densely pubescent to tomentose, abaxial usually whitish. | subsp. sericata |
1. Bracts (1–)2–10(–50) mm distant from sepals, margins entire, lobed, or toothed. | → 2 |
2. Stems erect, sometimes intertwined; leaf blades usually linear to narrowly triangular, sometimes ovate, base not lobed or hastate-lobed and lobes ± linear, oblong, or triangular, 1-pointed. | C. longipes |
2. Stems ascending, decumbent, procumbent, trailing, or twining-climbing; leaf blades oblong, oblong-ovate, orbiculate, ovate, broadly to narrowly triangular, triangular-hastate, or palmately 7–9-lobed, base usually lobed and lobes rounded or 1–3-pointed, base rarely cuneate or ± truncate. | → 3 |
3. Leaf blades oblong, oblong-ovate, orbiculate, or ovate, base lobed and lobes rounded or base cuneate to ± truncate. | C. felix |
3. Leaf blades usually broadly to narrowly triangular, ovate-triangular, or triangular-hastate (daggerlike), or palmately 7–9-lobed, sometimes ± reniform, base usually lobed and lobes rounded or 1–3-pointed, base rarely cuneate or ± truncate. | → 4 |
4. Herbage glabrous. | → 5 |
5. Leaf blades ± triangular to ovate-triangular; bracts linear, margins entire or proximally lobed or toothed. | C. purpurata |
5. Leaf blades narrowly triangular-hastate; bracts elliptic to broadly elliptic-oblong, margins entire. | C. peirsonii |
4. Herbage usually hairy, at least near leaf blade sinus and/or tip of peduncle, sometimes glabrate or glabrescent. | → 6 |
6. Leaf blades palmately 7–9-lobed. | C. stebbinsii |
6. Leaf blades not palmately lobed. | → 7 |
7. Bract margins entire. | → 8 |
8. Leaf blades ± triangular-hastate, middle lobe ± lance-linear; bracts 1–2 mm distant from sepals, linear, 5–16(–20) × 0.5–1.5 mm. | C. vanzuukiae |
8. Leaf blades ± triangular; bracts (1–)3–7 mm distant from sepals, linear to linear-oblong, 4–13(–18) × 1–4(–5) mm. | C. occidentalis |
7. Bract margins proximally lobed or toothed. | → 9 |
9. Herbage tomentose to villous; leaf blades ± broadly to narrowly triangular, basal lobes 2(–3)-pointed. | C. malacophylla |
9. Herbage glabrate or hairy; leaf blades narrowly triangular or triangular-hastate, basal lobes rounded or 1–2-pointed. | → 10 |
10. Bracts 1–2 mm distant from sepals, linear, 5–16(–20) × 0.5–1.5 mm. | C. vanzuukiae |
10. Bracts 2–12(–15) mm distant from sepals, lanceolate, linear, linear-oblong, oblanceolate, or narrowly to broadly triangular, 5–22(–30) × 2–4(–7) mm. | C. occidentalis |
1. Bracts immediately subtending, less than 1 mm from, sepals, margins entire. | → 11 |
11. Corollas horticultural doubles, limbs multilobed; stamens and ovaries absent. | C. pubescens |
11. Corollas not doubles, limbs weakly 5-lobed, 5-angled, or entire; stamens and ovaries present. | → 12 |
12. Bracts 1.5–3.5(–4) mm wide. | → 13 |
13. Herbage tomentellous, tomentose, or villous. | C. collina |
13. Herbage glabrous or ± hairy, not tomentellous, tomentose, or villous. | → 14 |
14. Leaf blades narrowly triangular-hastate, base lobed, lobes ± oblong to rhombic, 1-pointed. | C. peirsonii |
14. Leaf blades rounded-deltate to triangular-hastate, base cuneate or lobed, lobes not oblong to rhombic and 1-pointed. | → 15 |
15. Herbage glabrous; bract apices acute to obtuse. | C. atriplicifolia |
15. Herbage sparsely hairy; bract apices acute. | C. subacaulis |
12. Bracts 4–30 mm wide. | → 16 |
16. Perennials or subshrubs, rootstock woody. | C. macrostegia |
16. Perennials, rhizomatous. | → 17 |
17. Leaf blades ± reniform, ± fleshy; corollas pink. | C. soldanella |
17. Leaf blades not reniform, not fleshy; corollas usually pink, cream, or white, rarely purple. | → 18 |
18. Stems usually erect, procumbent, or twining-climbing, sometimes trailing or proximally erect and distally twining-climbing; mostly c, e North America. | → 19 |
19. Leaf blades triangular to triangular-hastate, base lobed, lobes 2(–3)-pointed; corollas usually pink, sometimes purple or white, 21–32(–35) mm. | C. hederacea |
19. Leaf blades ± cordate, elliptic, elliptic-ovate, linear, oblong, ovate, broadly to narrowly triangular, or triangular-hastate, base cuneate, rounded, or lobed, lobes obtuse, rounded, or 1–2-pointed; corollas pink or white, (20–)35–65(–70) mm. | → 20 |
20. Stems erect. | C. spithamaea |
20. Stems usually twining-climbing, sometimes trailing or proximally erect and distally twining-climbing. | → 21 |
21. Leaf blades elliptic-ovate, basal lobes obtuse or rounded, surfaces sometimes whitish tomentose. | C. catesbeiana |
21. Leaf blades ± cordate, linear, oblong, oblong-ovate, oval, ovate, triangular, or triangular-hastate, basal lobes usually 1–2-pointed, sometimes rounded or 1-pointed, surfaces not whitish. | → 22 |
22. Leaf blade basal sinuses usually acute to rounded, sometimes quadrate, rarely closed; bracts proximally flat or keeled, not or scarcely saccate, margins not or scarcely enfolding sepals, apices acute to subobtuse or truncate. | C. sepium |
22. Leaf blade basal sinuses ± quadrate to rounded; bracts proximally saccate, margins enfolding sepals, apices obtuse to truncate. | C. silvatica |
18. Stems usually ascending-decumbent, sometimes procumbent, suberect, trailing, or proximally erect and distally weakly twining-climbing, or almost absent; mostly w North America (C. macounii plains and west). | → 23 |
23. Herbage glabrous. | C. atriplicifolia |
23. Herbage moderately or sparsely hairy, puberulent, tomentellous, tomentose, or villous. | → 24 |
24. Herbage sparsely hairy, hairs appressed. | C. subacaulis |
24. Herbage moderately or sparsely hairy, puberulent, tomentellous, tomentose, or villous, hairs not appressed. | → 25 |
25. Leaf blade basal lobes ± rhombic, rounded; east of California. | C. macounii |
25. Leaf blade basal lobes 1–3-pointed; California. | → 26 |
26. Stems mostly to 60(–100) cm; leaves not in basal rosettes. | C. malacophylla |
26. Stems to (2–)50 cm or almost absent; leaves usually in basal rosettes. | → 27 |
27. Herbage tomentellous, tomentose, or villous; leaf blade margins ± undulate. | C. collina |
27. Herbage moderately to sparsely hairy; leaf blade margins not notably undulate. | C. subacaulis |
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