Flora of North America species comparison

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Caltha leptosepala

Ranunculaceae

broad-leaf caltha, broad-leaf marshmarigold, elkslip, mountain marsh-marigold, white marsh-marigold, White Mountain marsh-marigold

buttercup family, crowfoot family
Habit Herbs, sometimes woody or herbaceous climbers or low shrubs, perennial or annual, often rhizomatous.
Stems

leafless or with 1 leaf, erect.

unarmed.
Leaves

blade undivided or more commonly divided or compound, base cordate, sometimes truncate or cuneate, margins entire, toothed, or incised;

venation pinnate or palmate.
Basal leaves

blade oblong-ovate to orbiculate-reniform, largest 1.5-11.5(-15) × 1-13cm, margins entire or crenate to dentate.

Inflorescences

1-2(-4)-flowered.

terminal or axillary, racemes, cymes, umbels, panicles, or spikes, or flowers solitary, flowers pedicellate or sessile.
Flowers

15-40 mm diam.;

sepals white to yellow (abaxially bluish), 8.5-23 mm.

bisexual, sometimes unisexual, inconspicuous or showy, radially or bilaterally symmetric;

sepaloid bracteoles absent;

perianth hypogynous;

sepals usually imbricate, 3-6(-20), distinct, often petaloid and colored, occasionally spurred;

petals 0-26, distinct (connate in Consolida), plane, cup-shaped, funnel-shaped, or spurred, conspicuous or greatly reduced;

nectary usually present, rarely absent;

stamens 5-many, distinct;

anthers dehiscing longitudinally;

staminodes absent (except in Aquilegia and Clematis);

pistils 1-many;

styles present or absent, often persistent in fruit as beak.
Fruits

achenes, follicles, or rarely utricles, capsules, or berries, often aggregated into globose to cylindric heads.
Seeds

elliptic, 1.9-2.5 mm.

1-many per ovary, never stalked, not arillate;

endosperm abundant;

embryo usually small.
Follicles

4-15, spreading, short-stipitate or sessile, linear-oblong;

bodies 10-20 × 3-4.5 mm;

style and stigma straight or curved, 0.5-1.8 mm.

2n

=48,96.

Caltha leptosepala

Ranunculaceae
Phenology Flowering late spring–summer (Jun–Aug).
Habitat Open, wet, subalpine and alpine marshes, wet seepages
Elevation 750-3900 m (2500-12800 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; YT
[WildflowerSearch map]
[BONAP county map]
Worldwide
[BONAP county map]
Discussion

Caltha leptosepala is morphologically complex, and a number of segregate taxa have been described. Plants are most commonly assigned to two species, however. Caltha leptosepala in strict sense is found in the Rocky Mountains of Arizona and New Mexico north to Alaska and is characterized by longer-than-broad leaves with small, nonoverlapping basal lobes, solitary-flowered inflorescences, and sessile follicles. Plants in the Coast Ranges of central California north to the coastal islands of southern Alaska, distinguished by broader-than-long leaves with large, overlapping basal lobes, 2-flowered inflorescences, and stipitate follicles, have been called C.biflora. My comparison of specimens from the Rocky Mountains and the Coast Ranges indicated that no clear distinction could be made (table 1). While plants are often distinctive in the southern part of their range, a continuous intergradation between the two extremes exists over much of their range.

Table 1: [see original page on floranorthamerica.org]

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera ca. 60, species 1700 (22 genera, 284 species in the flora).

The flowers of many species of Ranunculaceae begin to open long before anthesis, while the floral organs are just partly expanded. Only mature flowers with open anthers should be used for determination of diagnostic characteristics (especially measurements).

The literature is inconsistent about the term for the whorl of organs between sepals and stamens; these may be conspicuous and petaloid, or reduced to stalked nectaries, or intermediate between the two states. They have been called petals, honey-leaves, or (when they are inconspicuous) staminodes or nectaries. We follow M. Tamura (1993) and treat as petals all organs between the sepals and stamens, except in Clematis and Aquilegia where they usually bear rudimentary anthers and clearly represent staminodes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Flowers bilaterally symmetric; sepals showy; petals smaller than sepals.
→ 2
1. Flowers radially symmetric; sepals showy or not; petals present or absent, smaller to larger than sepals.
→ 4
2. Upper (adaxial) sepal (hood) saccate or helmet-shaped; petals completely hidden by sepals.
Aconitum
2. Upper (adaxial) sepal spurred; petals at least partly exserted from calyx.
→ 3
3. Perennials; pistils 3(-5); petals 4, distinct.
Delphinium
3. Annuals; pistil 1; petals 2, connate.
Consolida
4. Fruits achenes or utricles; ovule 1 per pistil.
→ 5
4. Fruits follicles, capsules, or berries; ovules 2 or more per pistil (1 of 2 aborting in Xanthorhiza, leaving 1 seed at maturity).
→ 12
5. Sepals spurred; leaves all basal, blade linear or narrowly oblanceolate.
Myosurus
5. Sepals plane; leaves either not all basal, or blade not linear or narrowly oblanceolate.
→ 6
6. Leaves all cauline and opposite; stems ±woody, at least at base.
Clematis
6. Leaves cauline and alternate (rarely opposite), or basal, or plants with basal leaves and opposite or whorled involucral bracts; stems herbaceous.
→ 7
7. Plants with 1 or more pairs (opposite) or whorls of involucral bracts, these leaflike or calyxlike.
→ 8
7. Plants without involucral bracts (inconspicuous, linear-lanceolate involucral bracts in Trautvetteria), cauline leaves if present alternate (rarely a pair of opposite, unlobed leaves in Ranunculus sect. Flammula).
→ 9
8. Achenes with conspicuous veins or ribs on lateral surfaces; style absent.
T. thalictroides
8. Achenes without veins on lateral surfaces; style present.
Anemone
9. Petals absent; inflorescences panicles, racemes, or corymbs (umbels in Thalictrum thalictroides); filaments filiform or dilated distally.
→ 10
9. Petals present (rarely absent in Ranunculus pedatifidus); inflorescences simple or compound cymes or flowers solitary; filaments filiform.
→ 11
10. Leaves simple, blade lobed; flowers bisexual; inflorescences corymbs.
Trautvetteria
10. Leaves compound; flowers unisexual or bisexual; inflorescences panicles, racemes, corymbs, or umbels.
Thalictrum
11. Petals without nectaries; sepals 5(-8).
Adonis
11. Petals with basal nectaries; sepals 3-5(-6).
Ranunculus
12. Leaves dissected into linear, threadlike segments; pistils compound; fruits capsules.
Nigella
12. Leaves not dissected, if parted or compound the segments not linear; pistils simple; fruits aggregates of follicles or solitary or aggregate berries.
→ 13
13. Shrubs; beak of follicle lateral, strongly incurved against abaxial surface of follicle.
Xanthorhiza
13. Herbs; beak of follicle, if present, terminal or nearly so, straight or slightly curved, sometimes hooked at tip.
→ 14
14. Petals prominent, spurred.
Aquilegia
14. Petals if present inconspicuous, plane or funnel-shaped.
→ 15
15. Flowers 12-50, in racemes or racemelike panicles.
→ 16
15. Flowers 1-10, in leafy cymes or solitary.
→ 17
16. Pistils 1-8; fruits follicles, usually aggregate; petals 2-cleft or absent.
Cimicifuga
16. Pistil 1; fruits berries; petals unlobed.
Actaea
17. Leaves simple, blade often lobed 1/2-3/4 its length, margins entire, crenate, or toothed; petals absent.
→ 18
17. Leaves compound or divided to base; petals usually inconspicuous (absent in Enemion).
→ 19
18. Leaf blades unlobed, margins entire, dentate, or crenate; fruits follicles.
Caltha
18. Leaf blades lobed, margins serrate; fruits berries.
Hydrastis
19. Leaves ternately 1-2× compound.
→ 20
19. Leaves palmately or pedately compound or divided.
→ 21
20. Leaves all basal; leaf blade deeply divided, ternately or pinnately 1-2× compound; petals present.
Coptis
20. Leaves basal and cauline; leaf blade ternately 2× compound; petals absent.
Enemion
21. Leaf segments lobed, margins sharply toothed; sepals persistent in fruit.
Helleborus
21. Leaf segments cleft or parted, margins entire or toothed; sepals not persistent in fruit.
→ 22
22. Cauline leaves absent except for whorl of 3 involucral bracts immediately subtending flower; follicles stipitate.
Eranthis
22. Cauline leaves alternate, (0.8-)1 cm or more from flower, involucral whorl absent; follicles sessile.
Trollius
Source FNA vol. 3. FNA vol. 3, p. 85. Authors: Alan T. Whittemore, Bruce D. Parfitt.
Parent taxa Ranunculaceae > Caltha
Sibling taxa
C. natans, C. palustris
Subordinate taxa
Aconitum, Actaea, Adonis, Anemone, Aquilegia, Caltha, Cimicifuga, Clematis, Consolida, Coptis, Delphinium, Enemion, Eranthis, Helleborus, Hydrastis, Myosurus, Nigella, Ranunculus, Thalictrum, T. thalictroides, Trautvetteria, Trollius, Xanthorhiza
Synonyms C. biflora, C. biflora subsp. howellii, C. biflora var. rotundifolia, C. howellii, C. leptosepala var. rotundifolia, C. leptosepala var. sulfurea, C. uniflora, Psychropila leptosepala
Name authority de Candolle: Syst. Nat. 1: 310. (1817) Jussieu
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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