Calamagrostis stricta |
Poaceae subfam. pooideae |
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narrow hairgrass, narrowspike reedgrass, neglected reed grass, slim-stem reed grass, slipstem reed grass |
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Habit | Plants rarely with sterile culms; cespitose, usually with rhizomes shorter than 5 cm, 1-1.5 mm thick. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||||||
Culms | (10)35-90(120) cm, usually unbranched, smooth to slightly scabrous; nodes 1-3(4). |
usually hollow, sometimes solid. |
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Sheaths | usually smooth; collars usually smooth, sometimes scabrous, rarely pubescent; ligules (0.5)1-5.5(6) mm, truncate to obtuse, usually entire, sometimes lacerate; blades (5)11-25(34) cm long, (1)1.5-5(6) mm wide, flat or involute, usually scabrous, rarely smooth, sometimes puberulent. |
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Leaves | distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
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Panicles | (2)4-18(29) cm long, (0.7)1-2(2.8) cm wide, erect, contracted, sometimes interrupted, pale green to purple; branches 1.4-5(9.5) mm, smooth or scabrous, usually spikelet-bearing to or near the base, sometimes only to midlength. |
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Inflorescences | usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
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Spikelets | 2-4(5) mm; rachilla prolongations 0.5-1.5 mm, hairs 1.5-3 mm. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
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Glumes | usually less than 3 times as long as wide, rounded or keeled, usually smooth, rarely scabrous, keels smooth or scabrous, veins prominent to obscure, apices acute; callus hairs (1)1.5-3(4.5) mm, (0.5)0.7-0.9(1.3) times as long as the lemmas, abundant; lemmas 2-4(5) mm, 0.1-1.5 mm shorter than the glumes; awns 1.5-2.5 mm, usually attached to the lower 1/10-1/2 of the lemmas, rarely beyond the midpoint, equaling or exserted slightly beyond the margins of the glumes, usually stout, rarely slender, usually distinguishable from the callus hairs, straight or bent; anthers (0.9)1.2-1.8(2.4) mm, often sterile. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
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Caryopses | hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
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Calamagrostis stricta |
Poaceae subfam. pooideae |
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Distribution |
AK; AZ; CA; CO; CT; IA; ID; IL; IN; KS; MA; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; SD; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Greenland
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Discussion | Calamagrostis stricta grows throughout northern North America; it also is found in Europe and northeastern Asia. It grows in habitats ranging from meadows and grassland to wetlands, sandy shorelines, and sand dunes, from sea level to 3400 m. Primarily a species of open settings, it is frequently found in association with shrubs. Both subspecies have a notable but not exclusive association with alkaline to saline substrates. Calamagrostis stricta comprises both sexual and apomictic populations. Two subspecies, C. stricta subsp. stricta and subsp. inexpansa, intergrade but generally differ as described below. Greene (1984) treated subsp. inexpansa as consisting of the apomictic plants, probably derived from the sexual subsp. stricta. A number of apomictic variants were previously recognized at the species level; among these were C. lacustris (Kearney) Nash and C. fernaldii Louis-Marie, which are morphologically nearly indistinguishable from each other (Greene 1980, 1984). Plants of short stature and short inflorescences, growing in the north, have been referred to as Calamagrostis stricta subsp. borealis (C. Laest.) Á. Löve & D. Love or C. stricta var. borealis (C. Laest.) Hartm. These intergrade with taller plants; they are not recognized here as a distinct taxonomic entity. Calamagrostis stricta is sometimes confused with C. lapponica (see previous). In addition to the differences noted in the descriptions and key, the glumes of C. stricta are not as smooth and glossy, and are generally brown at the tip; those of C. lapponica are typically purple. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 729. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Calamagrostis | Poaceae | ||||||||||||||||||||||||||||||||||||||||
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Synonyms | C. stricta var. borealis, C. neglecta var. borealis, C. neglecta | |||||||||||||||||||||||||||||||||||||||||
Name authority | (Timm) Koeler | Benth. | ||||||||||||||||||||||||||||||||||||||||
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