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Hill-mustard, wartycabbage

crested wartycabbage

Habit Plants not scapose; glabrous or sparsely to densely pilose (multicellular glandular tubercles or papillae present throughout, except flowers). Annuals; glabrous or pubescent.
Stems

erect, often branched (many) distally.

unbranched or branched basally and distally, 1.5–7(–10) dm, (often white-hispid proximally and trichomes simple, glabrous distally).

Leaves

basal and cauline;

petiolate, sessile, or subsessile;

basal not rosulate [rosulate], petiolate, blade margins entire, pinnatifid, or lyrate;

cauline sessile (subsessile distally), blade (base cuneate, attenuate), margins dentate or entire.

Basal leaves

petiole 1–4(–6) cm;

blade lanceolate (in outline), lyrate to sinuate-pinnatifid, or runcinate-pinnatisect, (1–)3–15(–19) cm, lateral lobes oblong or lanceolate, (smaller than terminal lobe).

Cauline leaves

(distalmost) blade lanceolate or sublinear.

Racemes

(corymbose or paniculate), considerably elongated in fruit.

Flowers

sepals (yellowish green), oblong, (margins membranous), (glabrous, [pubescent or glandular]);

petals obovate, (longer than sepals), claw distinct or absent, (apex obtuse to emarginate);

stamens strongly tetradynamous;

filaments (yellowish), not dilated basally [dilated];

anthers oblong [ovate], (apex obtuse);

nectar glands confluent, subtending bases of stamens, median glands present.

sepals ascending to spreading, 3–4 × 1–1.5 mm;

petals (8–)10–13 × 4.5–6 mm, claw minute or absent;

filaments 3.5–6 mm;

anthers 1–1.3 mm.

Fruiting pedicels

divaricate, slender.

divaricate, straight, 1.5–2.5(–3) cm.

Fruits

nutletlike, sessile, (readily detached from pedicel), oblong, ovoid, or subglobose, smooth, terete, 4-angled, or with 4 cristate wings, (1–4-loculed), (woody);

valves (not distinct) not veined, glabrous;

replum not distinct;

septum subwoody or absent;

ovules 2–4 per ovary;

style obsolete or distinct, (slender, filiform or subconical);

stigma capitate.

oblong to subglobose, subquadrangular, with 4 irregularly dentate wings, 4-loculed, (0.8–)0.9–1.3(–1.4) cm × 5–6.5 mm;

style 3.5–5(–6) mm.

Seeds

plump [flattened], not winged, subglobose to ovoid [oblong];

seed coat (smooth), not mucilaginous when wetted;

cotyledons spirolobal.

3 or 4 per fruit, 2–3.5 mm.

x

= 7.

2n

= 14.

Bunias

Bunias erucago

Phenology Flowering Apr–Jun.
Habitat Roadsides, fields, pastures, disturbed areas, waste places
Distribution
from USDA
Europe; Asia; n Africa [Introduced in North America]
[BONAP county map]
from FNA
PA; VA; s Europe; sw Asia; n Africa [Introduced in North America]
[BONAP county map]
Discussion

Species 2 (2 in the flora).

Bunias erucago and B. orientalis have the same chromosome number; the former has only about 0.8-fold of the DNA amount of B. orientalis (J. Greilhuber and R. Obermayer 1999). Although both species are widespread weeds in Europe, they have not spread much in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits 4-winged, 4-loculed, (0.8-)0.9-1.3(-1.4) cm; seeds 3 or 4; petals (8-)10-13 mm; styles 3.5-5(-6) mm.
B. erucago
1. Fruits not winged, 1- or 2-loculed, (0.5-)0.6-0.7(-0.8) cm; seeds 1 or 2; petals (4-)4.5-7(-8) mm; styles obsolete or, rarely, to 1 mm.
B. orientalis
Source FNA vol. 7, p. 444. Author: Ihsan A. Al-Shehbaz. FNA vol. 7, p. 444.
Parent taxa Brassicaceae > tribe Buniadeae Brassicaceae > tribe Buniadeae > Bunias
Sibling taxa
B. orientalis
Subordinate taxa
B. erucago, B. orientalis
Name authority Linnaeus: Sp. Pl. 2: 669. (1753): Gen. Pl. ed. 5, 300. (1754) Linnaeus: Sp. Pl. 2: 670. (1753)
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