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Hill-mustard, wartycabbage

Habit Plants not scapose; glabrous or sparsely to densely pilose (multicellular glandular tubercles or papillae present throughout, except flowers). Annuals, biennials, or perennials; glandular (glands multicellular on multiseriate stalks).
Stems

erect, often branched (many) distally.

Leaves

basal and cauline;

petiolate, sessile, or subsessile;

basal not rosulate [rosulate], petiolate, blade margins entire, pinnatifid, or lyrate;

cauline sessile (subsessile distally), blade (base cuneate, attenuate), margins dentate or entire.

Cauline leaves

sessile or subsessile [petiolate];

blade base not auriculate, margins dentate or entire.

Trichomes

stalked, forked, or simple.

Racemes

(corymbose or paniculate), considerably elongated in fruit.

ebracteate, often elongated in fruit.

Flowers

sepals (yellowish green), oblong, (margins membranous), (glabrous, [pubescent or glandular]);

petals obovate, (longer than sepals), claw distinct or absent, (apex obtuse to emarginate);

stamens strongly tetradynamous;

filaments (yellowish), not dilated basally [dilated];

anthers oblong [ovate], (apex obtuse);

nectar glands confluent, subtending bases of stamens, median glands present.

actinomorphic;

sepals spreading or ascending [erect], lateral pair not saccate basally;

petals yellow, claw usually present [absent], often distinct;

filaments unappendaged [winged];

pollen 3-colpate.

Fruiting pedicels

divaricate, slender.

Fruits

nutletlike, sessile, (readily detached from pedicel), oblong, ovoid, or subglobose, smooth, terete, 4-angled, or with 4 cristate wings, (1–4-loculed), (woody);

valves (not distinct) not veined, glabrous;

replum not distinct;

septum subwoody or absent;

ovules 2–4 per ovary;

style obsolete or distinct, (slender, filiform or subconical);

stigma capitate.

silicles [siliques], indehiscent [dehiscent], unsegmented, terete or 4-angled [latiseptate];

ovules 2–4[–numerous] per ovary;

style obsolete or distinct;

stigma entire [2-lobed].

Seeds

plump [flattened], not winged, subglobose to ovoid [oblong];

seed coat (smooth), not mucilaginous when wetted;

cotyledons spirolobal.

aseriate [biseriate];

cotyledons spirolobal.

x

= 7.

Bunias

Brassicaceae tribe Buniadeae

Distribution
from USDA
Europe; Asia; n Africa [Introduced in North America]
[BONAP county map]
Europe; Asia; Africa [Introduced in North America]
Discussion

Species 2 (2 in the flora).

Bunias erucago and B. orientalis have the same chromosome number; the former has only about 0.8-fold of the DNA amount of B. orientalis (J. Greilhuber and R. Obermayer 1999). Although both species are widespread weeds in Europe, they have not spread much in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genus 1, species 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits 4-winged, 4-loculed, (0.8-)0.9-1.3(-1.4) cm; seeds 3 or 4; petals (8-)10-13 mm; styles 3.5-5(-6) mm.
B. erucago
1. Fruits not winged, 1- or 2-loculed, (0.5-)0.6-0.7(-0.8) cm; seeds 1 or 2; petals (4-)4.5-7(-8) mm; styles obsolete or, rarely, to 1 mm.
B. orientalis
Source FNA vol. 7, p. 444. Author: Ihsan A. Al-Shehbaz. FNA vol. 7, p. 443.
Parent taxa Brassicaceae > tribe Buniadeae Brassicaceae
Subordinate taxa
B. erucago, B. orientalis
Name authority Linnaeus: Sp. Pl. 2: 669. (1753): Gen. Pl. ed. 5, 300. (1754) de Candolle: Mém. Mus. Hist. Nat. 7: 245. (1821)
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