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bulbostyle, hairsedge

capillary hairsedge, Elliott's hairsedge

Habit Herbs, annual or perennial, cespitose, scapose, not rhizomatous. Herbs, annual or perennial, cespitose, slender.
Roots

diffuse, very fine.

Culms

stiff or flaccid, usually terete, ribbed.

to 40 cm, wiry.

Leaves

basal, polystichous, spreading to ascending;

sheaths open apically, apex fimbriate-ciliate, rarely entire;

ligules absent, lateral tufts of hair at junction of blade and sheath;

blades mostly linear or filiform, coarsely ribbed, usually longer than sheaths, 1 mm wide or less, mostly strongly involute, margins variably scabrid-ciliate.

¼–1/2 length of culms;

sheaths stramineous to tan or brown, glabrous or scabrid on ribs;

blades spreading to ascending, filiform, 0.5 mm wide, involute, margin glabrous or scabrid, abaxially glabrous or hirtellous.

Inflorescences

terminal on slender scapes, rarely axillary to culm leaves, simple or compound anthelae or spikelet solitary;

spikelets 1–50+;

involucral bracts (1–)2–8, spreading or erect, leaflike or scalelike.

terminal, in simple or compound anthelae;

scapes wiry, coarsely ribbed, glabrous or strumose-puberulent;

longer primary involucral bracts with setaceous blades exceeding or exceeded by inflorescence.

Spikelets

mostly ovoid to lanceoloid or lance-cylindric;

scales 6–50, spirally arranged, rarely nearly distichous, each subtending flower or 1–2 proximal scales empty.

red-brown to dark brown, lanceoloid to cylindric, longer than wide, 2–6 mm;

fertile scales broadly ovate, keeled, 1–1.3 mm, apex acute, abaxially puberulent, midrib included or excurrent as mucro.

Flowers

bisexual;

perianth absent;

stamens 1–3;

styles slender, (2–)3-fid, glabrous, base enlarged, persistent in fruit.

stamens 2–3;

anthers narrowly oblong, 0.8–1 mm.

Achenes

trigonous or 3-lobed, rarely biconvex, usually 3-ribbed.

mostly waxy gray, trigonous, obovoid, 0.8–1.5 mm, faces evenly papillate, finely rugulose or level;

tubercle a small, dark, compressed-conic button.

Bulbostylis

Bulbostylis ciliatifolia

Distribution
map from USDA
Mostly of dry or periodically dry; sunny; sandy uplands; particularly savanna in warm-temperate and tropical regions worldwide
[BONAP county map]
map from FNA
AL; FL; GA; LA; MS; NC; SC; TN; TX; VA; West Indies (Cuba)
[WildflowerSearch map]
[BONAP county map]
Discussion

Species ca. 100 (8 in the flora).

Culms of bulbostylises are coarse or fine, leafy only at the base, and terminating in scape (or rarely not scapose). Scapes are linear to filiform, wiry, variously ribbed. Spikelets or florets are short-stalked, even sessile in leaf axils. Flowers are protandrous and subsessile on short pedicellar joints; anthers are mostly linear-oblong or oblong-elliptic, two- to four-sporangiate, bilocular; style base is articulated to ovary summit and persists on fruit as buttonlike tubercle.

Etymology: Latin bulbus, bulb, and stylus, style

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Cyperaceae Cyperaceae > Bulbostylis
Sibling taxa
B. barbata, B. capillaris, B. funckii, B. juncoides, B. schaffneri, B. stenophylla, B. warei
Subordinate taxa
B. barbata, B. capillaris, B. ciliatifolia, B. funckii, B. juncoides, B. schaffneri, B. stenophylla, B. warei
B. ciliatifolia var. ciliatifolia, B. ciliatifolia var. coarctata
Key
1. Spikelets sessile or subsessile, forming globose to hemispheric head, head subtended by setaceous-tipped or cuspidate involucral bracts.
→ 2
1. Spikelets either solitary at culm tip or in anthelae, sometimes short-stalked (B. capillares), at plant base in leaf axils.
→ 4
2. Achene faces finely reticulate, not transversely rugose or rugulose; achenes 0.5–0.6 mm; spikelets red, red-brown, or dull brown; longer involucral bracts rarely exceeding inflorescence.
B. barbata
2. Achene faces transversely rugose or rugulose; achenes 1–1.2 mm; spikelets pale or greenish or tan; longer involucral bracts exceeding inflorescence.
→ 3
3. Involucral bracts with broad, scarious, strongly pectinate-fimbriate sheath border abruptly narrowing to blade; plants perennial, to 50 cm; anthers 3, 3 mm.
B. warei
3. Involucral bracts with scarious, entire border gradually narrowing to blade; plants annual, to 20 cm; anthers 1, 0.5 mm.
B. stenophylla
4. Spikelets 1 per culm; plants diminutive, delicate annuals.
→ 5
4. Spikelets prevalently more than 1 per culm, in anthelae; plants tall annuals or perennials.
→ 6
5. Spikelets of 2 sorts, sterile and fertile, respectively producing non-viable achenes, 1 mm or less, or viable achenes, 1.5 mm, in sheath axils at culm base.
B. funckii
5. Spikelets fertile, achenes all viable, 1 mm.
B. schaffneri
6. Surface of ripe achenes waxy gray, evenly papillate; Coastal Plain of United States.
B. ciliatifolia
6. Surface of ripe achenes pale red-brown, greenish or yellow-brown, tan, or gray, minute alveolae or papillae raised into finely or coarsely transversely rugose or rugulose faces; wider distribution.
→ 7
7. Plants hard-based, perennial; achene surfaces gray, yellow-brown or dark brown, rugulose, papillate; stamens 3; anthers 1 mm.
B. juncoides
7. Plants soft-based, annual; achene surfaces mostly yellow to pale brown, rugose, not papillate; stamens (1–)2; anthers 0.5–0.7 mm.
B. capillaris
1. Plants annual, 10–20(–30) cm; scapes mostly 0.5 mm diam.; anthelae mostly simple, open; longest involucral bract shorter than inflorescence; fertile scales dark red-brown or dull brown.
var. ciliatifolia
1. Plants perennial, 15–40 cm; scapes mostly 0.5–0.7 mm diam.; anthelae mostly compound, open or dense; proximalmost involucral bract longer than inflorescence; spikelets red brown or pale brown.
var. coarctata
Synonyms Oncostylis, Stenophyllus Scirpus ciliatifolius, Isolepis ciliatifolius, Stenophyllus ciliatifolius
Name authority Kunth: Enum. Pl. 2: 205. 1837, name conserved, not Steven 1817 or de Candolle (1836) (Elliott) Fernald: Rhodora 40: 391. (1938)
Source FNA vol. 23, p. 131. Treatment author: Robert Kral. FNA vol. 23, p. 134. Treatment author: Robert Kral.
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