Bromus tectorum |
Poaceae subfam. pooideae |
|||||||||||||||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
cheat brome, cheat grass, downy brome, downy chess, drooping brome |
|
|||||||||||||||||||||||||||||||||||||
Habit | Plants annual. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||
Culms | 5-90 cm, erect, slender, puberulent below the panicle. |
usually hollow, sometimes solid. |
||||||||||||||||||||||||||||||||||||
Sheaths | usually densely and softly retrorsely pubescent to pilose, upper sheaths sometimes glabrous; auricles absent; ligules 2-3 mm, glabrous, obtuse, lacerate; blades to 16 cm long, 1-6 mm wide, both surfaces softly hairy. |
|||||||||||||||||||||||||||||||||||||
Leaves | distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
|||||||||||||||||||||||||||||||||||||
Panicles | 5-20 cm long, 3-8 cm wide, open, lax, drooping distally, usually 1-sided; branches 1-4 cm, drooping, usually 1-sided and longer than the spikelets, usually at least 1 branch with 4-8 spikelets. |
|||||||||||||||||||||||||||||||||||||
Inflorescences | usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
|||||||||||||||||||||||||||||||||||||
Spikelets | 10-20 mm, usually shorter than the panicle branches, sides parallel or diverging distally, moderately laterally compressed, often purplish-tinged, not densely crowded, with 4-8 florets. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
||||||||||||||||||||||||||||||||||||
Glumes | villous, pubescent, or glabrous, margins hyaline; lower glumes 4-9 mm, 1-veined; upper glumes 7-13 mm, 3-5-veined; lemmas 9-12 mm, lanceolate, glabrous or pubescent to pilose, 5-7-veined, rounded over the midvein, margins hyaline, often with some hairs longer than those on the backs, apices acuminate, hyaline, bifid, teeth 0.8-2(3) mm; awns 10-18 mm, straight, arising 1.5 mm or more below the lemma apices; anthers 0.5-1 mm. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
||||||||||||||||||||||||||||||||||||
Caryopses | hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
|||||||||||||||||||||||||||||||||||||
2n | = 14. |
|||||||||||||||||||||||||||||||||||||
Bromus tectorum |
Poaceae subfam. pooideae |
|||||||||||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; AB; BC; MB; NB; NS; NT; ON; QC; SK; YT; Greenland
|
|||||||||||||||||||||||||||||||||||||
Discussion | Bromus tectorum is a European species that is well established in the Flora region and other parts of the world. It grows in disturbed sites, such as overgrazed rangelands, fields, sand dunes, road verges, and waste places. In the southwestern United States, Bromus tectorum is considered a good source of spring feed for cattle, at least until the awns mature. It is highly competitive and dominates rapidly after fire, especially in sagebrush areas. The resulting dense, fine fuels permanently shorten the fire-return interval, further hindering reestablishment of native species. It now dominates large areas of the sagebrush ecosystem of the western Flora region. Specimens with glabrous spikelets have been called Bromus tectorum f. nudus (Klett & Richt.) H. St. John. They occur throughout the range of the species, and are not known to have any other distinguishing characteristics. For this reason, they are not given formal recognition in this treatment. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||||||||||||||||||
Key |
|
|||||||||||||||||||||||||||||||||||||
Source | FNA vol. 24, p. 226. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Genea | Poaceae | ||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||
Synonyms | B. tectorum var. nudus, B. tectorum var. glabratus, Anisantha tectorum | |||||||||||||||||||||||||||||||||||||
Name authority | L. | Benth. | ||||||||||||||||||||||||||||||||||||
Web links |
|