Bromus hordeaceus |
Poaceae subfam. pooideae |
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brome mou, lesser soft brome, lopgrass, soft brome, soft chess |
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Habit | Plants annual or biennial. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||||||||||||||
Culms | 2-70 cm, erect or ascending. |
usually hollow, sometimes solid. |
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Leaves | distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
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Panicles | 1-13 cm long, 1-4 cm wide, erect, usually ovoid, open, becoming dense, occasionally reduced to 1 or 2 spikelets; branches shorter than the spikelets, ascending to erect, straight or almost so. |
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Inflorescences | usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
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Spikelets | (11)14-20(23) mm, lanceolate, terete to moderately laterally compressed; florets 5-10, bases concealed at maturity; rachilla internodes concealed at maturity. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
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Glumes | pilose or glabrous; lower glumes 5-7 mm, 3-5-veined; upper glumes 6.5-8 mm, 5-7-veined; lemmas 6.5-11 mm long, 3-5 mm wide, lanceolate, chartaceous, antrorsely pilose to pubescent, or glabrous proximally or throughout, 7-9-veined, lateral veins prominently ribbed, rounded over the midvein, hyaline margins abruptly or bluntly angled, not inrolled at maturity, apices acute, bifid, teeth shorter than 1 mm; awns 6-8 mm, usually arising less than 1.5 mm below the lemma apices, straight to recurved at maturity; anthers 0.6-1.5 mm. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
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Caryopses | equaling or shorter than the paleas, thin, weakly inrolled to flat. |
hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
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Lower | sheaths densely, often retrorsely, pilose; upper sheaths pubescent or glabrous; ligules 1-1.5 mm, hairy, obtuse, erose; blades 2-19 cm long, 1-4 mm wide, abaxial surfaces glabrous or hairy, adaxial surfaces hairy. |
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2n | = 28. |
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Bromus hordeaceus |
Poaceae subfam. pooideae |
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Distribution |
AK; AR; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; WA; WI; WY; HI; AB; BC; LB; NB; NS; NT; ON; QC; YT; Greenland
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Discussion | Bromus hordeaceus is native to southern Europe and northern Africa. It is weedy, growing in disturbed areas such as roadsides, fields, sandy beaches, and waste places, and can be found in many locations in the Flora region, with the exception of the central Canadian provinces and most of the southeastern United States. Its origin is obscure. Ainouche et al. (1999) reviewed various suggestions, and concluded that at least one of its diploid ancestors may have been an extinct or undiscovered species related to B. caroli-henrici, a diploid species. The four subspecies are usually morphologically distinct. Ainouche et al. (1999), however, found no evidence of genetic differentiation among them. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 232. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Bromus | Poaceae | ||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | B. mollis | |||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | L. | Benth. | ||||||||||||||||||||||||||||||||||||||||||||||||
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