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brome mou, lesser soft brome, lopgrass, soft brome, soft chess

meadow brome

Habit Plants annual or biennial. Plants perennial; cespitose, shortly rhizomatous.
Culms

2-70 cm, erect or ascending.

30-90 cm, erect or decumbent, forming distinct clumps;

nodes 2-3, glabrous or puberulent;

internodes glabrous or puberulent.

Sheaths

glabrous or with hairs;

auricles to 1 mm on the lower leaves;

ligules 0.4-1.0 mm, glabrous or ciliate, truncate, erose;

blades 10-20 cm long, 2-3 mm wide, scabridulous, glabrous or sparsely pilose, margins sometimes ciliate.

Panicles

1-13 cm long, 1-4 cm wide, erect, usually ovoid, open, becoming dense, occasionally reduced to 1 or 2 spikelets;

branches shorter than the spikelets, ascending to erect, straight or almost so.

8-20 cm long, lax;

branches scabridulous, with 1-2 spikelets.

Spikelets

(11)14-20(23) mm, lanceolate, terete to moderately laterally compressed;

florets 5-10, bases concealed at maturity;

rachilla internodes concealed at maturity.

20-32 mm, lanceolate, becoming cuneate, with 5-8 florets.

Glumes

pilose or glabrous;

lower glumes 5-7 mm, 3-5-veined;

upper glumes 6.5-8 mm, 5-7-veined;

lemmas 6.5-11 mm long, 3-5 mm wide, lanceolate, chartaceous, antrorsely pilose to pubescent, or glabrous proximally or throughout, 7-9-veined, lateral veins prominently ribbed, rounded over the midvein, hyaline margins abruptly or bluntly angled, not inrolled at maturity, apices acute, bifid, teeth shorter than 1 mm;

awns 6-8 mm, usually arising less than 1.5 mm below the lemma apices, straight to recurved at maturity;

anthers 0.6-1.5 mm.

glabrous, sometimes scabridulous on the veins;

lower glumes 6.5-10 mm, 1(3)-veined;

upper glumes 7.5-12 mm, 3-5-veined;

lemmas 10-13 mm, oblong to lanceolate, rounded over the midvein, 7-veined, glabrous or appressed-hairy, sometimes scabridulous, apices acute, entire or minutely bifid;

awns 4-8 mm, straight or slightly spreading, arising less than 1.5 mm below the lemma apices;

anthers 2.5-5.2 mm.

Caryopses

equaling or shorter than the paleas, thin, weakly inrolled to flat.

Lower

sheaths densely, often retrorsely, pilose;

upper sheaths pubescent or glabrous;

ligules 1-1.5 mm, hairy, obtuse, erose;

blades 2-19 cm long, 1-4 mm wide, abaxial surfaces glabrous or hairy, adaxial surfaces hairy.

2n

= 28.

= 70.

Bromus hordeaceus

Bromus riparius

Distribution
from FNA
AK; AR; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; WA; WI; WY; HI; AB; BC; LB; NB; NS; NT; ON; QC; YT; Greenland
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Discussion

Bromus hordeaceus is native to southern Europe and northern Africa. It is weedy, growing in disturbed areas such as roadsides, fields, sandy beaches, and waste places, and can be found in many locations in the Flora region, with the exception of the central Canadian provinces and most of the southeastern United States. Its origin is obscure. Ainouche et al. (1999) reviewed various suggestions, and concluded that at least one of its diploid ancestors may have been an extinct or undiscovered species related to B. caroli-henrici, a diploid species.

The four subspecies are usually morphologically distinct. Ainouche et al. (1999), however, found no evidence of genetic differentiation among them.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Bromus riparius is an Asian species that was introduced to the United States in the late 1950s for cultivation as a pasture grass. Various cultivars are now grown, mainly in Canada and the northwestern United States. The description given here is derived in part from cultivated specimens. North American plants have sometimes been referred to, incorrectly, as Bromus biebersteinii Roem. & Schult. (Vogel et al. 1996). Bromus riparius differs from that species in having acute lemma apices and, usually, more pubescent leaf blades, sheaths, and lemmas.

The existence of Bromus riparius in the Flora region was not realized until shortly before this treatment was submitted for publication, making it impossible to fully investigate its similarities to B. inermis and B. pumpellianus, particularly subsp. dicksonii. It appears to differ from both species in having shorter culms on average, longer awns than B. inermis, and shorter rhizomes than B. pumpellianus subsp. pumpellianus. Its distribution in the Flora region is not known.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lemmas (7)8-11 mm long, usually pubescent or pilose.
→ 2
2. Awns more than 0.1 mm wide at the base, straight, erect; culms (3)10-70 cm long
subsp. hordeaceus
2. Awns less than 0.1 mm wide at the base, often divaricate or recurved at maturity; culms 15-25(60) cm long
subsp. molliformis
1. Lemmas 6.5-8(9) mm long, glabrous or pubescent.
→ 3
3. Culms (3)10-70 cm long; panicles up to 10 cm long, usually with more than 1 spikelet; lemmas usually glabrous; caryopses usually as long as the paleas; habitat various
subsp. pseudothomineii
3. Culms 2-16 cm long; panicles 1-3 cm long, often reduced to 1 spikelet; lemmas pubescent or glabrous; caryopses shorter than the paleas; plants of maritime or lacustrine sands
subsp. thotninei
Source FNA vol. 24, p. 232. FNA vol. 24, p. 206.
Parent taxa Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Bromus Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Bromopsis
Sibling taxa
B. aleutensis, B. anomalus, B. arenarius, B. arizonicus, B. arvensis, B. berteroanus, B. briziformis, B. carinatus, B. caroli-henrici, B. catharticus, B. ciliatus, B. commutatus, B. danthoniae, B. diandrus, B. erectus, B. frondosus, B. grandis, B. hallii, B. inermis, B. japonicus, B. kalmii, B. laevipes, B. lanatipes, B. lanceolatus, B. latiglumis, B. lepidus, B. madritensis, B. maritimus, B. mucroglumis, B. nottowayanus, B. orcuttianus, B. pacificus, B. polyanthus, B. porteri, B. pseudolaevipes, B. pubescens, B. pumpellianus, B. racemosus, B. ramosus, B. richardsonii, B. riparius, B. rubens, B. scoparius, B. secalinus, B. sitchensis, B. squarrosus, B. sterilis, B. suksdorfii, B. tectorum, B. texensis, B. vulgaris
B. aleutensis, B. anomalus, B. arenarius, B. arizonicus, B. arvensis, B. berteroanus, B. briziformis, B. carinatus, B. caroli-henrici, B. catharticus, B. ciliatus, B. commutatus, B. danthoniae, B. diandrus, B. erectus, B. frondosus, B. grandis, B. hallii, B. hordeaceus, B. inermis, B. japonicus, B. kalmii, B. laevipes, B. lanatipes, B. lanceolatus, B. latiglumis, B. lepidus, B. madritensis, B. maritimus, B. mucroglumis, B. nottowayanus, B. orcuttianus, B. pacificus, B. polyanthus, B. porteri, B. pseudolaevipes, B. pubescens, B. pumpellianus, B. racemosus, B. ramosus, B. richardsonii, B. rubens, B. scoparius, B. secalinus, B. sitchensis, B. squarrosus, B. sterilis, B. suksdorfii, B. tectorum, B. texensis, B. vulgaris
Subordinate taxa
B. hordeaceus subsp. hordeaceus, B. hordeaceus subsp. molliformis, B. hordeaceus subsp. pseudothomineii, B. hordeaceus subsp. thotninei
Synonyms B. mollis
Name authority L. Rehmann
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