Bromus hordeaceus |
Bromus pumpellianus |
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brome mou, lesser soft brome, lopgrass, soft brome, soft chess |
arctic brome, pumpelly brome |
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Habit | Plants annual or biennial. | Plants perennial; usually rhizomatous, sometimes cespitose, rhizomes short to long-creeping. | ||||||||||||||||
Culms | 2-70 cm, erect or ascending. |
50-135 cm, erect or ascending, sometimes geniculate, usually single or few together, sometimes clumped; nodes 2-7, pubescent or glabrous; internodes glabrous or pubescent. |
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Sheaths | pilose, villous, or glabrous; auricles sometimes present on the lower leaves; ligules to 4 mm, glabrous, truncate or obtuse, erose; blades 7-30 cm long, 2.5-8.5(9) mm wide, flat, pubescent or glabrous on both surfaces, sometimes only the adaxial surface pubescent. |
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Panicles | 1-13 cm long, 1-4 cm wide, erect, usually ovoid, open, becoming dense, occasionally reduced to 1 or 2 spikelets; branches shorter than the spikelets, ascending to erect, straight or almost so. |
10-24 cm, open or contracted, erect or nodding; branches erect to spreading. |
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Spikelets | (11)14-20(23) mm, lanceolate, terete to moderately laterally compressed; florets 5-10, bases concealed at maturity; rachilla internodes concealed at maturity. |
16-32(45) mm, elliptic to lanceolate, terete to moderately laterally compressed, sometimes purplish, with 4-14 florets. |
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Glumes | pilose or glabrous; lower glumes 5-7 mm, 3-5-veined; upper glumes 6.5-8 mm, 5-7-veined; lemmas 6.5-11 mm long, 3-5 mm wide, lanceolate, chartaceous, antrorsely pilose to pubescent, or glabrous proximally or throughout, 7-9-veined, lateral veins prominently ribbed, rounded over the midvein, hyaline margins abruptly or bluntly angled, not inrolled at maturity, apices acute, bifid, teeth shorter than 1 mm; awns 6-8 mm, usually arising less than 1.5 mm below the lemma apices, straight to recurved at maturity; anthers 0.6-1.5 mm. |
glabrous or hairy; lower glumes (4)5-10 mm, 1(3)-veined; upper glumes (5)7.5-13 mm, 3-veined; lemmas 9-16 mm, lanceolate, rounded over the midvein, sparsely to densely hairy throughout, or on the margins and lower portion of the back, or along the marginal veins and keel, apices subulate to acute, entire or slightly emarginate, lobes shorter than 1 mm; awns usually present, sometimes absent, to 7.5 mm, straight, arising less than 1.5 mm below the lemma apices; anthers 3.5-7 mm. |
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Caryopses | equaling or shorter than the paleas, thin, weakly inrolled to flat. |
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Lower | sheaths densely, often retrorsely, pilose; upper sheaths pubescent or glabrous; ligules 1-1.5 mm, hairy, obtuse, erose; blades 2-19 cm long, 1-4 mm wide, abaxial surfaces glabrous or hairy, adaxial surfaces hairy. |
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2n | = 28. |
= 28, 56. |
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Bromus hordeaceus |
Bromus pumpellianus |
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Distribution |
AK; AR; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; WA; WI; WY; HI; AB; BC; LB; NB; NS; NT; ON; QC; YT; Greenland
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Discussion | Bromus hordeaceus is native to southern Europe and northern Africa. It is weedy, growing in disturbed areas such as roadsides, fields, sandy beaches, and waste places, and can be found in many locations in the Flora region, with the exception of the central Canadian provinces and most of the southeastern United States. Its origin is obscure. Ainouche et al. (1999) reviewed various suggestions, and concluded that at least one of its diploid ancestors may have been an extinct or undiscovered species related to B. caroli-henrici, a diploid species. The four subspecies are usually morphologically distinct. Ainouche et al. (1999), however, found no evidence of genetic differentiation among them. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The range of Bromus pumpellianus extends from Asia to North America, where it includes Alaska, the western half of Canada, the western United States as far south as New Mexico, and a few other locations eastward. It is sometimes treated as a subspecies of B. inermis. It differs from that species primarily in its tendency to have pubescent lemmas, nodes, and leaf blades. Two subspecies that differ in morphology and distribution are described below. Both strongly resemble the recently introduced Bromus riparius, differing in the case of B. pumpellianus subsp. pumpellianus in having longer rhizomes, or, in the case of B. pumpellianus subsp. dicksonii, in having a more restricted distribution. It is possible that the description and distribution of B. pumpellianus may be based in part on misidentification of B. riparius, as many taxonomists may have been unaware of the introduction of the latter species to North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 232. | FNA vol. 24, p. 207. | ||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Bromus | Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Bromopsis | ||||||||||||||||
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Synonyms | B. mollis | B. pumpellianus var. arcticus, B. inermis subsp. pumpellianus, B. inermis var. purpurascens, Bromopsis pumpelliana var. arctica, Bromopsis pumpelliana, Bromopsis inermis subsp. pumpellianus | ||||||||||||||||
Name authority | L. | Scribn. | ||||||||||||||||
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