Bromus hordeaceus |
Poaceae tribe Bromeae |
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brome mou, lesser soft brome, lopgrass, soft brome, soft chess |
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Habit | Plants annual or biennial. | Plants annual or perennial; usually cespitose, sometimes rhizomatous. | ||||||||||||
Culms | 2-70 cm, erect or ascending. |
annual, not woody, not branching above the base; internodes usually hollow, rarely solid. |
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Sheaths | closed, margins united for most of their length; collars without tufts of hair on the sides; auricles sometimes present; ligules membranous, sometimes shortly ciliate, those of the upper and lower cauline leaves usually similar; pseudopetioles absent; blades linear to narrowly lanceolate, venation parallel, cross venation not evident, without arm or fusoid cells, cross sections non-Kranz, epidermes without microhairs, not papillate. |
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Panicles | 1-13 cm long, 1-4 cm wide, erect, usually ovoid, open, becoming dense, occasionally reduced to 1 or 2 spikelets; branches shorter than the spikelets, ascending to erect, straight or almost so. |
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Inflorescences | usually terminal panicles, sometimes reduced to racemes in depauperate plants; disarticulation above the glumes and beneath each floret. |
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Spikelets | (11)14-20(23) mm, lanceolate, terete to moderately laterally compressed; florets 5-10, bases concealed at maturity; rachilla internodes concealed at maturity. |
5-80 mm, not viviparous, terete to laterally compressed, with 3-30 bisexual florets, distal florets sometimes reduced; rachillas prolonged beyond the bases of the distal florets. |
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Glumes | pilose or glabrous; lower glumes 5-7 mm, 3-5-veined; upper glumes 6.5-8 mm, 5-7-veined; lemmas 6.5-11 mm long, 3-5 mm wide, lanceolate, chartaceous, antrorsely pilose to pubescent, or glabrous proximally or throughout, 7-9-veined, lateral veins prominently ribbed, rounded over the midvein, hyaline margins abruptly or bluntly angled, not inrolled at maturity, apices acute, bifid, teeth shorter than 1 mm; awns 6-8 mm, usually arising less than 1.5 mm below the lemma apices, straight to recurved at maturity; anthers 0.6-1.5 mm. |
usually unequal, rarely more or less equal, exceeded by the distal florets, usually longer than 1/4 the length of the adjacent florets, lanceolate, 1-9(11)-veined; florets terete to laterally compressed; calluses glabrous, not well developed; lemmas lanceolate to ovate, rounded or keeled over the midvein, herbaceous to coriaceous, 5-13-veined, veins converging somewhat distally, apices usually minutely bilobed to bifid, rarely entire, usually awned, sometimes unawned, awns unbranched, terminal or subterminal, usually straight, sometimes geniculate; paleas usually shorter than the lemmas; lodicules 2, glabrous, not veined; anthers 3; ovaries with hairy apices; styles 2, bases free. |
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Caryopses | equaling or shorter than the paleas, thin, weakly inrolled to flat. |
narrowly ellipsoid to linear, longitudinally grooved; hila linear; embryos about 1/6 the length of the caryopses. |
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Lower | sheaths densely, often retrorsely, pilose; upper sheaths pubescent or glabrous; ligules 1-1.5 mm, hairy, obtuse, erose; blades 2-19 cm long, 1-4 mm wide, abaxial surfaces glabrous or hairy, adaxial surfaces hairy. |
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x | = 7. |
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2n | = 28. |
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Bromus hordeaceus |
Poaceae tribe Bromeae |
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Distribution |
AK; AR; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; WA; WI; WY; HI; AB; BC; LB; NB; NS; NT; ON; QC; YT; Greenland
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Discussion | Bromus hordeaceus is native to southern Europe and northern Africa. It is weedy, growing in disturbed areas such as roadsides, fields, sandy beaches, and waste places, and can be found in many locations in the Flora region, with the exception of the central Canadian provinces and most of the southeastern United States. Its origin is obscure. Ainouche et al. (1999) reviewed various suggestions, and concluded that at least one of its diploid ancestors may have been an extinct or undiscovered species related to B. caroli-henrici, a diploid species. The four subspecies are usually morphologically distinct. Ainouche et al. (1999), however, found no evidence of genetic differentiation among them. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
There are three genera in the Bromeae. One genus, Bromus, grows in the Flora region. The tribe was included in the Festuceae Dumort. [= Poeae] by earlier agrostologists (e.g., Hitchcock 1951) because it has paniculate inflorescences, spikelets with more than 1 floret, and glumes that are shorter than the lemmas. It is now considered to be most closely related to the Triticeae. This is indicated by the pubescent apices of the ovaries and simple endosperm starch grains. It is further supported by data from serology, nucleic acid sequences, and seedling development. These data do not support a close relationship between the Bromeae and Brachypodium, a genus that has sometimes been included in the tribe. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 232. | FNA vol. 24, p. 192. | ||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Bromus | Poaceae > subfam. Pooideae | ||||||||||||
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Synonyms | B. mollis | |||||||||||||
Name authority | L. | Dumort. | ||||||||||||
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