Bromus catharticus |
Poaceae subfam. pooideae |
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rescue brome, rescue grass, rescuegras |
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Habit | Plants annual, biennial, or perennial; loosely cespitose or tufted. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||||||
Culms | 30-120 cm tall, 2-4 mm thick, erect or decumbent. |
usually hollow, sometimes solid. |
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Sheaths | usually densely, often retrorsely, hairy, hairs sometimes confined to the throat; auricles absent; ligules 1-4 mm, glabrous or pilose, obtuse, lacerate to erose; blades 4-30 cm long, 3-10 mm wide, flat, glabrous or hairy on both surfaces. |
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Leaves | distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
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Panicles | 9-28 cm, usually open, erect or nodding; lower branches shorter than 10 cm, 1-4 per node, spreading or ascending, with up to 5 spikelets variously distributed. |
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Inflorescences | usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
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Spikelets | (17)20-40 mm, shorter than at least some pedicels and branches, elliptic to lanceolate, strongly laterally compressed, not crowded or overlapping, with 4-12 florets. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
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Glumes | smooth or scabrous, glabrous or pubescent; lower glumes 7-12 mm, 5-7(9)-veined; upper glumes 9-17 mm, 7-9(11)-veined, shorter than the lowest lemma; lemmas 11-20 mm, lanceolate, laterally compressed, strongly keeled, usually glabrous, sometimes pubescent distally, smooth or scabrous, 9-13-veined, veins often raised and riblike, margins sometimes conspicuous, hyaline, whitish or partly purplish, apices entire or toothed, teeth acute, shorter than 1 mm; awns absent or to 10 mm; anthers 0.5-1 mm in cleistogamous florets, 2-5 mm in chasmogamous florets. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
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Caryopses | hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
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2n | = 42. |
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Bromus catharticus |
Poaceae subfam. pooideae |
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Distribution |
AL; AR; AZ; CA; CO; DC; FL; GA; IA; IL; KS; KY; LA; MD; MO; MS; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SC; SD; TN; TX; UT; VA; HI; AB; NF; ON
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Discussion | The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 199. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Ceratochloa | Poaceae | ||||||||||||||||||||||||||||||||||||||||
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Synonyms | Ceratochloa unioloides, B. willdenowii, B. unioloides | |||||||||||||||||||||||||||||||||||||||||
Name authority | Vahl | Benth. | ||||||||||||||||||||||||||||||||||||||||
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