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Arizona brome

great brome, ripgut brome, ripgut grass

Habit Plants annual; tufted. Plants annual.
Culms

30-90 cm tall, to 3 mm thick, erect.

20-90 cm, erect or decumbent, puberulent below the panicle.

Sheaths

retrorsely pilose, sometimes mostly glabrous, throats sometimes with hairs;

auricles absent;

ligules 1-4 mm, usually glabrous, obtuse, erose;

blades 8-18 cm long, 3-9 mm wide, flat, sparsely pilose on both surfaces or the abaxial surfaces glabrous.

softly pilose, hairs often retrorse or spreading;

auricles absent;

ligules 2-3 mm, glabrous, obtuse, lacerate or erose;

blades 3.5-27 cm long, 1-9 mm wide, both surfaces pilose.

Panicles

12-25 cm, somewhat contracted or open;

lower branches shorter than 10 cm, 2-3(5) per node, initially erect to ascending, spreading at maturity, with 1-2 spikelets variously distributed.

13-25 cm long, 2-12 cm wide, erect to spreading;

branches 1-7 cm, stiffly erect to ascending or spreading, with 1 or 2 spikelets.

Spikelets

18-25 mm, elliptic to lanceolate, strongly laterally compressed, with 4-8 florets.

25-70 mm, sides parallel or diverging distally, moderately laterally compressed, with 4-11 florets.

Glumes

subequal, smooth or scabrous;

lower glumes 8-12.5 mm, 3-veined;

upper glumes 9.5-14 mm, 7-veined, about as long as the lowest lemma;

lemmas 9.5-14 mm, lanceolate, laterally compressed, prominently 7-veined, strongly keeled at least distally, glabrous or pubescent distally or throughout, marginal hairs, if present, longer than those elsewhere, apices entire or with acute teeth shorter than 1 mm;

awns 6-13 mm, sometimes slightly geniculate;

anthers 0.4-0.5 mm.

smooth or scabrous, margins hyaline;

lower glumes 15-25 mm, 1-3-veined;

upper glumes 20-35 mm, 3-5-veined;

lemmas 20-35 mm, linear-lanceolate, scabrous, 7-veined, rounded over the midvein, margins hyaline, apices bifid, acuminate, teeth 3-5 mm;

awns 30-65 mm, straight, arising 1.5 mm or more below the lemma apices;

anthers 0.5-1 mm.

2n

= 84.

= 42, 56.

Bromus arizonicus

Bromus diandrus

Distribution
from FNA
AZ; CA; NV; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; AZ; CA; CO; DC; DE; GA; ID; IL; LA; MA; MD; MO; MT; NC; NJ; NM; NV; NY; OK; OR; SC; TX; UT; VA; WA; HI; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Bromus arizonicus grows in dry, open areas and disturbed ground of the southwest, usually below 2000 m. Its range extends from California and southern Nevada into Arizona, New Mexico, and northern Mexico.

Stebbins et al. (1944) demonstrated that, like Bromus carinatus var. carinatus, B. arizonicus obtained three of its genomes from B. catharticus or a close relative, but the remaining three genomes are not homologous with those in B. carinatus, probably being derived from a species in a section other than Ceratochloa. The small anthers of B. arizonicus strongly suggest that most seed is produced by selfing.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Bromus diandrus is native to southern and western Europe. It is now established in North America, where it grows in disturbed ground, waste places, fields, sand dunes, and limestone areas. It occurs from southwestern British Columbia to Baja California, Mexico, and eastward to Montana, Colorado, Texas, and scattered locations in the eastern United States. The common name 'ripgut grass' indicates the effect it has on animals if they consume the sharp, long-awned florets of this species.

Bromus diandrus, as treated here, includes B. rigidus Roth. Sales (1993) reduced these two taxa to varietal rank, pointing out that the differences between them in panicle morphology and callus and scar shape are subtle enough that identification of many specimens beyond B. diandrus sensu lato is often impossible.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 201. FNA vol. 24, p. 224.
Parent taxa Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Ceratochloa Poaceae > subfam. Pooideae > tribe Bromeae > Bromus > sect. Genea
Sibling taxa
B. aleutensis, B. anomalus, B. arenarius, B. arvensis, B. berteroanus, B. briziformis, B. carinatus, B. caroli-henrici, B. catharticus, B. ciliatus, B. commutatus, B. danthoniae, B. diandrus, B. erectus, B. frondosus, B. grandis, B. hallii, B. hordeaceus, B. inermis, B. japonicus, B. kalmii, B. laevipes, B. lanatipes, B. lanceolatus, B. latiglumis, B. lepidus, B. madritensis, B. maritimus, B. mucroglumis, B. nottowayanus, B. orcuttianus, B. pacificus, B. polyanthus, B. porteri, B. pseudolaevipes, B. pubescens, B. pumpellianus, B. racemosus, B. ramosus, B. richardsonii, B. riparius, B. rubens, B. scoparius, B. secalinus, B. sitchensis, B. squarrosus, B. sterilis, B. suksdorfii, B. tectorum, B. texensis, B. vulgaris
B. aleutensis, B. anomalus, B. arenarius, B. arizonicus, B. arvensis, B. berteroanus, B. briziformis, B. carinatus, B. caroli-henrici, B. catharticus, B. ciliatus, B. commutatus, B. danthoniae, B. erectus, B. frondosus, B. grandis, B. hallii, B. hordeaceus, B. inermis, B. japonicus, B. kalmii, B. laevipes, B. lanatipes, B. lanceolatus, B. latiglumis, B. lepidus, B. madritensis, B. maritimus, B. mucroglumis, B. nottowayanus, B. orcuttianus, B. pacificus, B. polyanthus, B. porteri, B. pseudolaevipes, B. pubescens, B. pumpellianus, B. racemosus, B. ramosus, B. richardsonii, B. riparius, B. rubens, B. scoparius, B. secalinus, B. sitchensis, B. squarrosus, B. sterilis, B. suksdorfii, B. tectorum, B. texensis, B. vulgaris
Synonyms B. berteroanus var. excelsus B. rigidus var. gussonei, B. rigidus, Anisantha diandra
Name authority (Shear) Stebbins Roth
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