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Coulter's brickellbush

false boneset

Habit Shrubs, 30–150 cm. Perennials, 30–200 cm (bases woody).
Stems

much branched from bases, pubescent, often gland-dotted.

branched, pubescent.

Leaves

opposite;

petioles 4–12 mm;

blades 3-nerved from bases, broadly ovate to deltate, 10–60 × 10–30 mm, bases truncate, hastate, or subcordate, margins usually toothed (1–3 sets of sharp teeth near bases), rarely subentire, apices attenuate, faces minutely pubescent.

mostly opposite (alternate in vars. gracillima and texana);

petioles 0–10 mm;

blades 1- or 3-nerved from bases, lanceolate, lance-linear, lance-ovate, lance-rhombic, linear, or oblong, 25–100 × 0.5–40 mm, bases acute, margins entire or ± dentate (often revolute), apices obtuse to acuminate, faces glandular-pubescent.

Peduncles

5–40 mm, pubescent and sometimes gland-dotted.

5–100 mm, glandular-pubescent.

Involucres

cylindric to campanulate, 8–12 mm.

cylindric to narrowly campanulate, 7–15 mm.

Florets

13–25;

corollas pale yellow-green, often purple-tinged, 7–8.2 mm.

6–35;

corollas pale yellow, yellow-green, pinkish lavender, or maroon, 4.5–6 mm.

Phyllaries

17–22 in 4–6 series, greenish, often purple-tinged, 4–5-striate, unequal, margins scarious (apices acute to acuminate or mucronate, minutely pubescent);

outer lance-ovate to narrowly lanceolate (glabrous or sparsely pubescent), inner narrowly lanceolate to linear (glabrous).

22–26 in 4–6 series, green to stramineous, sometimes purple-tinged, 3–7-striate, unequal, margins scarious (often ciliate);

outer ovate to lance-ovate (puberulent, often densely gland-dotted, apices acute to acuminate), inner lanceolate (± gland-dotted, apices obtuse to aristate).

Heads

in loose, paniculiform arrays.

in paniculiform or corymbiform arrays.

Cypselae

3–5 mm, hispidulous, strigose, or glandular-pubescent;

pappi of 28–40 smooth or barbellulate bristles.

2.7–5.5 mm, glabrous or strigose, sometimes hispidulous or velutinous and/or gland-dotted;

pappi of 20–28 white or tawny, usually plumose or subplumose, sometimes barbellate, bristles.

Brickellia coulteri

Brickellia eupatorioides

Distribution
from FNA
AZ; NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CO; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; MT; NC; ND; NE; NJ; NM; OH; OK; PA; SC; SD; TN; TX; UT; VA; WI; WV; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (2 in the flora}.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 6 (6 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Midstem leaf blades mostly 40–60 mm; peduncles mostly 15–40 mm, mostly glandular-pubescent
var. coulteri
1. Midstem leaf blades mostly 20–40(–50) mm; peduncles 5–25 mm, pubescent, mostly lacking glandular hairs
var. brachiata
1. Leaf blades lance-linear or linear, 0.5–3 mm wide
→ 2
1. Leaf blades lanceolate, lance-linear, lance-ovate, or lance-rhombic, 1–40 mm wide
→ 4
2. Florets 9–13
var. floridana
2. Florets 16–30
→ 3
3. Leaf blades 1–3 mm wide (lengths of proximal leaves greater than distals); involucres 8–13 mm
var. chlorolepis
3. Leaf blades 0.5–1 mm wide (lengths of proximal leaves equaling distals); involucres 5–10 mm
var. gracillima
4. Phyllaries: lengths of outers and mids 1/2–1 times inners, apices long-acuminate, usuallycontorted
var. texana
4. Phyllaries: lengths of outers and mids to 1/2 times inners, apices obtuse, acute, or acuminate, not contorted
→ 5
5. Leaves 1-nerved from bases; petioles 0–1 mm; heads borne singly or 2–3 in loose, paniculiform arrays
var. chlorolepis
5. Leaves 1- or 3-nerved from bases; petioles 0–10 mm; heads mostly 3–8 in dense, corymbiform arrays.
→ 6
6. Involucres 7–11 mm; florets 6–15
var. eupatorioides
6. Involucres 8–15 mm; florets 15–35
var. corymbulosa
Source FNA vol. 21, p. 496. FNA vol. 21, p. 498.
Parent taxa Asteraceae > tribe Eupatorieae > Brickellia Asteraceae > tribe Eupatorieae > Brickellia
Sibling taxa
B. amplexicaulis, B. atractyloides, B. baccharidea, B. betonicifolia, B. brachyphylla, B. californica, B. chenopodina, B. cordifolia, B. cylindracea, B. dentata, B. desertorum, B. eupatorioides, B. floribunda, B. frutescens, B. grandiflora, B. greenei, B. hinckleyi, B. incana, B. knappiana, B. laciniata, B. lemmonii, B. longifolia, B. microphylla, B. nevinii, B. oblongifolia, B. parvula, B. pringlei, B. rusbyi, B. simplex, B. venosa, B. veronicifolia
B. amplexicaulis, B. atractyloides, B. baccharidea, B. betonicifolia, B. brachyphylla, B. californica, B. chenopodina, B. cordifolia, B. coulteri, B. cylindracea, B. dentata, B. desertorum, B. floribunda, B. frutescens, B. grandiflora, B. greenei, B. hinckleyi, B. incana, B. knappiana, B. laciniata, B. lemmonii, B. longifolia, B. microphylla, B. nevinii, B. oblongifolia, B. parvula, B. pringlei, B. rusbyi, B. simplex, B. venosa, B. veronicifolia
Subordinate taxa
B. coulteri var. brachiata, B. coulteri var. coulteri
B. eupatorioides var. chlorolepis, B. eupatorioides var. corymbulosa, B. eupatorioides var. eupatorioides, B. eupatorioides var. floridana, B. eupatorioides var. gracillima, B. eupatorioides var. texana
Synonyms Kuhnia eupatorioides
Name authority A. Gray: Smithsonian Contr. Knowl. 3(5): 86. (1852) (Linnaeus) Shinners: Sida 4: 274. (1971)
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