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Coulter's brickellbush

gravelbar brickellbush

Habit Shrubs, 30–150 cm. Perennials, 40–120 cm (caudices woody).
Stems

much branched from bases, pubescent, often gland-dotted.

branched, pubescent, gland-dotted.

Leaves

opposite;

petioles 4–12 mm;

blades 3-nerved from bases, broadly ovate to deltate, 10–60 × 10–30 mm, bases truncate, hastate, or subcordate, margins usually toothed (1–3 sets of sharp teeth near bases), rarely subentire, apices attenuate, faces minutely pubescent.

mostly opposite, sometimes alternate or subopposite;

petioles 0–3 mm;

blades 3-nerved from bases, lanceolate to obovate, 20–60 × 12–30 mm, bases ± cuneate, margins crenate to serrate, apices obtuse to acute, abaxial faces tomentose, often densely gland-dotted as well (venation raised, reticulate), adaxial faces pubescent and gland-dotted.

Peduncles

5–40 mm, pubescent and sometimes gland-dotted.

2–15 mm, densely tomentose, often gland-dotted.

Involucres

cylindric to campanulate, 8–12 mm.

cylindric to campanulate, 8–13 mm.

Florets

13–25;

corollas pale yellow-green, often purple-tinged, 7–8.2 mm.

10–21;

corollas greenish, yellow, or yellow-orange, often purple-tinged, 5–7 mm.

Phyllaries

17–22 in 4–6 series, greenish, often purple-tinged, 4–5-striate, unequal, margins scarious (apices acute to acuminate or mucronate, minutely pubescent);

outer lance-ovate to narrowly lanceolate (glabrous or sparsely pubescent), inner narrowly lanceolate to linear (glabrous).

18–22 in 5–7 series, greenish, often purplish-tinged, 6–8-striate, unequal, margins scarious (apices acute to acuminate);

outer ovate (sparsely villous, gland-dotted, lengths no more than 1/2 inner, margins long-ciliate), inner narrowly lanceolate to linear-oblong (glabrous).

Heads

in loose, paniculiform arrays.

in ± dense, racemiform or paniculiform arrays.

Cypselae

3–5 mm, hispidulous, strigose, or glandular-pubescent;

pappi of 28–40 smooth or barbellulate bristles.

3.5–5 mm, sparsely to densely pubescent;

pappi of 30–35, white to tawny, barbellate bristles.

2n

= 18.

Brickellia coulteri

Brickellia cylindracea

Phenology Flowering Jun–Nov.
Habitat Dry limestone hillsides
Elevation 100–600 m (300–2000 ft)
Distribution
from FNA
AZ; NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Varieties 3 (2 in the flora}.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

L. D. Flyr (1970) rightly noted that Brickellia cylindracea is variable in leaf size, shape, and texture as well as arrangements of heads, and added that it is not easily distinguished from B. conduplicata (= B. lemmonii var. conduplicata), which accounts for most of the specimens in west Texas. Brickellia cylindracea has a more eastern distribution than B. lemmonii var. conduplicata and appears to have a strong preference for limestone soils. Some specimens from limestone outcrops in west Texas blur distinction between the two species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Midstem leaf blades mostly 40–60 mm; peduncles mostly 15–40 mm, mostly glandular-pubescent
var. coulteri
1. Midstem leaf blades mostly 20–40(–50) mm; peduncles 5–25 mm, pubescent, mostly lacking glandular hairs
var. brachiata
Source FNA vol. 21, p. 496. FNA vol. 21, p. 497.
Parent taxa Asteraceae > tribe Eupatorieae > Brickellia Asteraceae > tribe Eupatorieae > Brickellia
Sibling taxa
B. amplexicaulis, B. atractyloides, B. baccharidea, B. betonicifolia, B. brachyphylla, B. californica, B. chenopodina, B. cordifolia, B. cylindracea, B. dentata, B. desertorum, B. eupatorioides, B. floribunda, B. frutescens, B. grandiflora, B. greenei, B. hinckleyi, B. incana, B. knappiana, B. laciniata, B. lemmonii, B. longifolia, B. microphylla, B. nevinii, B. oblongifolia, B. parvula, B. pringlei, B. rusbyi, B. simplex, B. venosa, B. veronicifolia
B. amplexicaulis, B. atractyloides, B. baccharidea, B. betonicifolia, B. brachyphylla, B. californica, B. chenopodina, B. cordifolia, B. coulteri, B. dentata, B. desertorum, B. eupatorioides, B. floribunda, B. frutescens, B. grandiflora, B. greenei, B. hinckleyi, B. incana, B. knappiana, B. laciniata, B. lemmonii, B. longifolia, B. microphylla, B. nevinii, B. oblongifolia, B. parvula, B. pringlei, B. rusbyi, B. simplex, B. venosa, B. veronicifolia
Subordinate taxa
B. coulteri var. brachiata, B. coulteri var. coulteri
Name authority A. Gray: Smithsonian Contr. Knowl. 3(5): 86. (1852) A. Gray & Engelmann: Proc. Amer. Acad. Arts 1: 46. (1847)
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