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needle grama

red grama, three-awn grama

Habit Plants annual; tufted. Plants perennial; cespitose, older plants occasionally shortly rhizomatous.
Culms

4-60 cm, outer culms of a tuft decumbent, sometimes geniculate, branched at the lower nodes.

5-40 cm, slender, wiry, erect or slightly geniculate at the lower nodes;

lower internodes glabrous, shorter than those above.

Leaves

mostly basal;

sheaths glabrous, sometimes scabridu-lous, becoming flattened, persistent;

ligules 0.2-0.5 mm, of hairs;

blades 0.7-8 cm long, 0.5-1.5(2) mm wide, scabridulous, margins often with papillose-based hairs basally.

Panicles

2.5-10.5 cm, with (1)4-15 branches;

branches 5-45 mm, deciduous, densely pubescent (at least basally), with 2-10 spikelets per branch, axes extending 2-10 mm beyond the base of the terminal spikelets, apices entire;

disarticulation at the base of the branches, the break forming a sharp tip.

3-9 cm, with 2-7 branches;

branches 7-25 mm, persistent, spreading, ascending, or appressed, straight to slightly arcuate, with 8-24(32) spikelets, axes terminating in a spikelet;

disarticulation above the glumes.

Spikelets

appressed.

appressed to pectinate, reddish-purple;

with 1 bisexual floret and 1 rudimentary floret.

Glumes

bilobed;

lower glumes 1.7-3.4 mm, slightly shorter than the upper glumes, veins excurrent to 0.6 mm;

upper glumes 1.9-4 mm, glabrous or pubescent, hairs not papillose-based, veins excurrent to 1 mm;

lower lemmas 1.2-2.2 mm, glabrous, sparsely appressed pubescent along the veins or densely appressed pubescent for much of their length and on the margins, trilobed, lobes veined, tapering into 3 awns, awns 2.2-6.6 mm, central awns not flanked by membranous lobes;

anthers 0.2-0.4 mm, yellow;

rachilla segments glabrous;

upper florets glabrous, of 3 equal awns, awns 2-7 mm.

Caryopses

2.5-3 mm.

0.8-1.5 mm long, 0.3-0.6 mm wide.

Ligules

0.2-0.5 mm, membranous, lacerate or ciliate;

blades 2-5(9) cm long, 0.7-2 mm wide, flat or folded, adaxial surfaces sometimes with papillose-based hairs, margins usually with papillose-based hairs near the ligules.

Proximal

spikelet on each branch with 1 floret;

lower glumes 1.5-3.5 mm, glabrous, narrow to subulate;

upper glumes 5.5-6.2 mm, densely pubescent, at least on the basal 1/2;

lemmas 5.8-6 mm, acuminate, unawned;

lowest paleas almost as long as the lemmas, bifid, glabrous;

rachillas prolonged beyond the florets for about 0.5 mm.

Distal

spikelets with 1 bisexual and 1 rudimentary floret, glumes unequal, glabrous, minutely scabrous on the keels; narrowly acute or acuminate;

lower glumes 1.5-2 mm;

upper glumes 5-6 mm, glabrous or sparsely pubescent basally, often divergent;

lowest lemmas 6-8 mm, veins pubescent, lateral veins excurrent as short (to 1 mm) awns, acuminate, midvein extended into a setaceous tip or a short awn;

lowest paleas 5-7 mm, bifid, veins often excurrent as short awns;

anthers about 2.5 mm, yellow or yellow and red;

distal florets reduced to a pubescent, 3-awned, awn column, awns 2-7 mm, exserted.

2n

= 40.

= 20.

Bouteloua aristidoides

Bouteloua trifida

Distribution
from FNA
AZ; CA; MD; NM; NV; TX; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; NV; TX; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

There are two varieties, both of which grow in the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Bouteloua trifida grows on dry open plains, shrubby hills, and rocky slopes, at 2200-2500 m. Its range extends from the southwestern United States to central Mexico. It is a drought-resistant species that is sometimes mistaken for Aristida because of its delicate, cespitose growth habit and purplish, 3-awned spikelets. Juvenile plants may also be confused with B. barbata but that species is annual, with the central awn flanked by two membranous lobes and the lowest paleas 4-lobed and 2-awned.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Panicle branches with 2-5 spikelets, usually 5-16 mm to the base of the terminal spikelets, axes usually extending an additional 6-10 mm
var. aristidoides
1. Panicle branches with 6-10 spikelets, usually 15-35 mm to the base of the terminal spikelets, axes extending an additional 2-5(7) mm
var. arizonica
1. Lower lemmas densely appressed pubescent; awns 2.2-4.5 mm long; anthers 0.2-0.3 mm long
var. burkii
1. Lower lemmas glabrous or sparsely appressed pubescent along both sides of the veins; awns (3.2)4-6.6 mm long; anthers 0.3-0.4 mm long
var. trifida
Source FNA vol. 25, p. 255. FNA vol. 25, p. 264.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Bouteloua > subg. Bouteloua Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Bouteloua > subg. Chondrosum
Sibling taxa
B. barbata, B. breviseta, B. chondrosoides, B. curtipendula, B. eludens, B. eriopoda, B. gracilis, B. hirsuta, B. kayi, B. parryi, B. radicosa, B. ramosa, B. repens, B. rigidiseta, B. simplex, B. trifida, B. uniflora, B. warnockii
B. aristidoides, B. barbata, B. breviseta, B. chondrosoides, B. curtipendula, B. eludens, B. eriopoda, B. gracilis, B. hirsuta, B. kayi, B. parryi, B. radicosa, B. ramosa, B. repens, B. rigidiseta, B. simplex, B. uniflora, B. warnockii
Subordinate taxa
B. aristidoides var. aristidoides, B. aristidoides var. arizonica
B. trifida var. burkii, B. trifida var. trifida
Name authority (Kunth) Griseb. Thurb. ex S. Watson
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