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botryche de virginie, common grapefern, rattlesnake fern

botryche lunaire, common moonwort, moonwort grapefern

Pinnae

to 12 pairs, usually approximate to overlapping, slightly ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, lanceolate, divided to tip.

to 9 pairs, spreading, mostly overlapping except in shaded forest forms, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, broadly fan-shaped, undivided to tip, margins mainly entire or undulate, rarely dentate, apical lobe usually cuneate to spatulate, notched, approximate to adjacent lobes, apex rounded, venation like ribs of fan, midribs absent.

Trophophore

sessile;

blade pale green, 3–4-pinnate, to 25 × 33 cm, thin, herbaceous.

stalk 0–1 mm;

blade dark green, oblong, 1-pinnate, to 10 × 4 cm, thick, fleshy.

Pinnules

lanceolate and deeply lobed, lobes linear, serrate, apex pointed, venation pinnate, midrib present.

Sporophores

2-pinnate, 0.5–1.5(–2) times length of trophophore.

1–2-pinnate, 0.8–2 times length of trophophore.

2n

=184.

=90.

Botrychium virginianum

Botrychium lunaria

Phenology Leaves seasonal, appearing in early spring and dying in late summer. Leaves appearing in spring, dying in latter half of summer.
Habitat Common to abundant, especially in shaded forests and shrubby second growth, rare or absent in arid regions Open fields, occasionally forests in southern occurrences
Elevation 0–1500 m (0–4900 ft) 0–3700 m (0–12100 ft)
Distribution
from FNA
AK; AL; AR; AZ; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Mexico; Central America; South America in Brazil; Colombia; Ecuador; Peru; Eurasia
from FNA
AK; AZ; CA; CO; ID; MA; ME; MI; MN; MT; NH; NM; NV; NY; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; SPM; Pacific Islands in New Zealand; Greenland; Eurasia; s South America; Australia
[WildflowerSearch map]
[BONAP county map]
Discussion

Botrychium virginianum is the most widespread Botrychium in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium lunaria grows with many other species of Botrychium, occasionally hybridizing with them. This species, geographically the most widespread of the moonworts, has notably uniform morphology.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Osmundopteris Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Synonyms Osmunda virginiana Osmunda lunaria, B. onondagense
Name authority (Linnaeus) Swartz: J. Bot. (Schrader) 1800(2): 111. (1801) (Linnaeus) Swartz: J. Bot. (Schrader) 1800(2): 110. (1801)
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