Botrychium virginianum |
Botrychium biternatum |
|
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botryche de virginie, common grapefern, rattlesnake fern |
southern grapefern, sparse-lobed grapefern, sparselobe grapefern |
|
Pinnae | to 12 pairs, usually approximate to overlapping, slightly ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, lanceolate, divided to tip. |
to 7 pairs, usually remote, horizontal, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, undivided except in proximal 2/3–1/2. |
Trophophore | sessile; blade pale green, 3–4-pinnate, to 25 × 33 cm, thin, herbaceous. |
stalk 4–20 cm, 2–2.5 times length of trophophore rachis; blade green to dark green, plane, 2–3-pinnate, to 18 × 28 cm, herbaceous. |
Pinnules | lanceolate and deeply lobed, lobes linear, serrate, apex pointed, venation pinnate, midrib present. |
elongate, obliquely lanceolate to narrowly lanceolate, margins nearly parallel and finely denticulate, apex short-acuminate, venation pinnate. |
Sporophores | 2-pinnate, 0.5–1.5(–2) times length of trophophore. |
1–2-pinnate, 2–3 times length of trophophore. |
2n | =184. |
=90. |
Botrychium virginianum |
Botrychium biternatum |
|
Phenology | Leaves seasonal, appearing in early spring and dying in late summer. | Leaves green over winter, sporophores seasonal, new leaves appearing in late spring–early summer. |
Habitat | Common to abundant, especially in shaded forests and shrubby second growth, rare or absent in arid regions | Frequent in low woods and brushy fields |
Elevation | 0–1500 m (0–4900 ft) | 0–600 m (0–2000 ft) |
Distribution |
AK; AL; AR; AZ; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Mexico; Central America; South America in Brazil; Colombia; Ecuador; Peru; Eurasia |
AL; AR; DE; FL; GA; IL; IN; KY; LA; MD; MO; MS; NC; NJ; OH; OK; PA; SC; TN; TX; VA; WV
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Discussion | Botrychium virginianum is the most widespread Botrychium in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Botrychium biternatum often grows with B. dissectum and B. jenmanii. The name B. biternatum was misapplied by L.Underwood to B. lunarioides (W.H. Wagner Jr. 1961). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Osmundopteris | Ophioglossaceae > Botrychium > subg. Sceptridium > sect. Sceptridium |
Sibling taxa | ||
Synonyms | Osmunda virginiana | Osmunda biternata, B. dissectum var. tenuifolium, B. tenuifolium |
Name authority | (Linnaeus) Swartz: J. Bot. (Schrader) 1800(2): 111. (1801) | (Savigny) L. Underwood: Bot. Gaz. 22: 407. (1896) |
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