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blunt-lobe grape-fern, blunt-lobed grapefern, botryche d'oneida, botryche du lac onéida

Crater Lake grapefern, pumice moonwort

Pinnae

to 5 pairs, usually remote, horizontal to ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, undivided except in proximal 2/3–3/4.

to 6 pairs, overlapping, strongly ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, asymmetrically cuneate, basal pinna pair often divided into 2 unequal parts, lobed to tip, margins entire, sinuate to shallowly crenate, apex rounded to truncate, venation pinnate.

Trophophore

stalk 2–15 cm, 1.5–2.5 times length of blade rachis;

blade dull bluish green, ± plane, 2–3-pinnate, to 15 × 20 cm, ± leathery.

stalk 0–10 mm, 0.1–0.5 times length of trophophore rachis;

blade dull, glaucous, whitish green, deltate, 2-pinnate, 4 × 6 cm, thickly leathery.

Pinnules

obliquely ovate, margins finely crenulate to denticulate, apex rounded to acute, venation pinnate.

Sporophores

2–3-pinnate, 1.5–2.5 times length of trophophore.

1–3-pinnate, 1–1.5 times length of trophophore.

2n

=90.

=90.

Botrychium oneidense

Botrychium pumicola

Phenology Leaves green over winter, sporophores seasonal, new leaves appearing in spring. Leaves appearing in summer.
Habitat In moist, shady, acidic woods and swamps Pumice scree
Elevation 0–1200 m (0–3900 ft) 1900–2500m (6200–8200ft)
Distribution
from FNA
CT; DC; DE; IN; KY; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; VA; VT; WI; WV; NB; ON; QC
[WildflowerSearch map]
from FNA
OR
[BONAP county map]
Discussion

Botrychium oneidense commonly occurs with B. dissectum and B. multifidum. Young individuals of both may resemble B. oneidense (W.H. Wagner Jr. 1961b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium pumicola is a famous narrow endemic known from only a few colonies on fully exposed pumice scree on the sides of and in the general vicinity of Crater Lake, Klamath and Deschutes counties, Oregon. This plant has a very congested appearance with an extremely compact sporangial cluster and overlapping pinnae. Like most other members of subg. Botrychium, the trophophore is located high on the common stalk, but the common stalk is subterranean, giving the impression that the leaf originates near ground level. Botrychium pumicola has been found growing with B. lanceolatum and B. simplex.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Sceptridium > sect. Sceptridium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Synonyms B. ternatum var. oneidense, B. dissectum var. oneidense, B. multifidum var. oneidense
Name authority (Gilbert) House: Amer. Midl. Naturalist 7: 126. (1905) Coville: in L. Underwood, Native Ferns ed. 6 69. (1900)
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