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blunt-lobe grape-fern, blunt-lobed grapefern, botryche d'oneida, botryche du lac onéida

botryche lunaire, common moonwort, moonwort grapefern

Pinnae

to 5 pairs, usually remote, horizontal to ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, undivided except in proximal 2/3–3/4.

to 9 pairs, spreading, mostly overlapping except in shaded forest forms, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, broadly fan-shaped, undivided to tip, margins mainly entire or undulate, rarely dentate, apical lobe usually cuneate to spatulate, notched, approximate to adjacent lobes, apex rounded, venation like ribs of fan, midribs absent.

Trophophore

stalk 2–15 cm, 1.5–2.5 times length of blade rachis;

blade dull bluish green, ± plane, 2–3-pinnate, to 15 × 20 cm, ± leathery.

stalk 0–1 mm;

blade dark green, oblong, 1-pinnate, to 10 × 4 cm, thick, fleshy.

Pinnules

obliquely ovate, margins finely crenulate to denticulate, apex rounded to acute, venation pinnate.

Sporophores

2–3-pinnate, 1.5–2.5 times length of trophophore.

1–2-pinnate, 0.8–2 times length of trophophore.

2n

=90.

=90.

Botrychium oneidense

Botrychium lunaria

Phenology Leaves green over winter, sporophores seasonal, new leaves appearing in spring. Leaves appearing in spring, dying in latter half of summer.
Habitat In moist, shady, acidic woods and swamps Open fields, occasionally forests in southern occurrences
Elevation 0–1200 m (0–3900 ft) 0–3700 m (0–12100 ft)
Distribution
from FNA
CT; DC; DE; IN; KY; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; VA; VT; WI; WV; NB; ON; QC
[WildflowerSearch map]
from FNA
AK; AZ; CA; CO; ID; MA; ME; MI; MN; MT; NH; NM; NV; NY; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; SPM; Pacific Islands in New Zealand; Greenland; Eurasia; s South America; Australia
[WildflowerSearch map]
[BONAP county map]
Discussion

Botrychium oneidense commonly occurs with B. dissectum and B. multifidum. Young individuals of both may resemble B. oneidense (W.H. Wagner Jr. 1961b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium lunaria grows with many other species of Botrychium, occasionally hybridizing with them. This species, geographically the most widespread of the moonworts, has notably uniform morphology.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Sceptridium > sect. Sceptridium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Synonyms B. ternatum var. oneidense, B. dissectum var. oneidense, B. multifidum var. oneidense Osmunda lunaria, B. onondagense
Name authority (Gilbert) House: Amer. Midl. Naturalist 7: 126. (1905) (Linnaeus) Swartz: J. Bot. (Schrader) 1800(2): 110. (1801)
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