Botrychium lanceolatum |
Ophioglossaceae |
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botryche élancé, lance-leaf grapefern, lance-leaf moonwort, red triangle moonwort, triangle moonwort |
Adder's-tongue family |
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Habit | Plants perennials, terrestrial or epiphytic. | |||||||||||||
Roots | lacking root hairs, unbranched or with a few narrow lateral branches, in 1 species dichotomously branched. |
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Stems | simple, unbranched, upright, with eustelic vascular tissue. |
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Leaves | bases dilated, clasping, forming sheath, open or fused, surrounding successive leaf primordia; primordia glabrous or with long, uniseriate hairs. |
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Pinnae | to 5 pairs, ascending, approximate, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, linear to broadly lanceolate, entire to divided to tip, margins with distinct lobes or segments, apex acute to rounded, venation pinnate. |
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Sporangia | exposed or embedded, 0.5–1.5 mm diam., thick-walled, with thousands of spores. |
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Spores | all 1 kind, trilete, thick-walled, surface rugate, tuberculate, baculate (with projecting rods usually higher than wide), sometimes joined in delicate network, mostly with ± warty surface. |
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Gametophytes | not green, usually fleshy, round or linear, subterranean, mycorrhizal. |
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Trophophore | stalk 0–1 mm; blade dull to shiny green to dark green, deltate, 1–2-pinnate, to 6 × 7 cm. |
blades compound to simple, rarely absent, veins anastomosing or free, pinnate, or arranged like ribs of fan. |
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Sporophores | 1–3-pinnate, 1–2.5 times length of trophophore, divided into several equally long branches (all other botrychiums have a single stalk or 1 dominant and 2 smaller). |
pinnately branched or simple. |
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Indument | absent or of widely scattered, long, uniseriate hairs, especially on petioles and rachises. |
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2n | =90. |
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Botrychium lanceolatum |
Ophioglossaceae |
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Distribution |
AK; AZ; CO; CT; ID; KY; MA; ME; MI; MN; MT; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; ON; QC; SK; YT; Eurasia
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Nearly worldwide |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Ophioglossaceae comprise two clearly defined subfamilies, Botrychioideae and Ophioglossoideae, which are sometimes recognized as distinct families. Ophioglossaceae may be only distantly related to the ferns and more closely related to Marattiales and certain seedplants, especially Cycadales, in such characteristics as stelar type, cork cambium, dilated leaf bases, conduplicate vernation, intercalary leaf growth, collateral leaf traces, circular-bordered pits, eusporangia, massive gametophytes, sunken archegonia, and presence in some species of endoscopic embryos. (Key to genera of Ophioglossaceae) Genera 5, species ca. 70–80 (3 genera, 38 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2, p. 85. | ||||||||||||
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | |||||||||||||
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Synonyms | Osmunda lanceolata | |||||||||||||
Name authority | (S. G. Gmelin) Angström: Bot. Not. 1854: 68. (1854) | Agardh | ||||||||||||
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