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botryche élancé, lance-leaf grapefern, lance-leaf moonwort, red triangle moonwort, triangle moonwort

botryche à segments spatulés, spathulate botrychium, spatulate moonwort, spoon-shape moonwort

Pinnae

to 5 pairs, ascending, approximate, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, linear to broadly lanceolate, entire to divided to tip, margins with distinct lobes or segments, apex acute to rounded, venation pinnate.

to 8 pairs, ascending, remote, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, mostly narrowly spatulate to linear-spatulate and rounded or ± 2-cleft, lobed to unlobed to tip, margins mainly entire or occasionally irregularly and shallowly incised, apex rounded-notched, venation like ribs of fan, midrib absent.

Trophophore

stalk 0–1 mm;

blade dull to shiny green to dark green, deltate, 1–2-pinnate, to 6 × 7 cm.

stalk 0–1 mm;

blade shiny yellow-green, narrowly deltate, flat, 1-pinnate, to 8 × 2.5 cm, thick, leathery.

Sporophores

1–3-pinnate, 1–2.5 times length of trophophore, divided into several equally long branches (all other botrychiums have a single stalk or 1 dominant and 2 smaller).

1–2-pinnate, 1.2–2 times length of trophophore.

2n

=90.

=180.

Botrychium lanceolatum

Botrychium spathulatum

Phenology Leaves appearing late spring through summer.
Habitat Sand dunes, old fields, and grassy railroad sidings
Elevation 0–2000 m (0–6600 ft)
Distribution
from FNA
AK; AZ; CO; CT; ID; KY; MA; ME; MI; MN; MT; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; ON; QC; SK; YT; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; MI; MT; AB; BC; NB; NT; ON; PE; QC; YT
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium spathulatum has long been confused with the more common B. minganense, with which it often grows in the Lake Superior region. The leaves appear later in B. spathulatum than in B. minganense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trophophore blade green to pale yellow-green, broad, coarse, succulent; middle and terminal segments usually more than 2 mm wide exclusive of lobes.
subsp. lanceolatum
1. Trophophore blade dark green, slender, delicate but firm; middle segments usually less than 2 mm wide exclusive of lobes.
subsp. angustisegmentum
Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. virginianum
Subordinate taxa
B. lanceolatum subsp. angustisegmentum, B. lanceolatum subsp. lanceolatum
Synonyms Osmunda lanceolata
Name authority (S. G. Gmelin) Angström: Bot. Not. 1854: 68. (1854) W. H. Wagner: Amer. Fern J. 80: 77. (1990)
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