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botryche élancé, lance-leaf grapefern, lance-leaf moonwort, red triangle moonwort, triangle moonwort

Crater Lake grapefern, pumice moonwort

Pinnae

to 5 pairs, ascending, approximate, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, linear to broadly lanceolate, entire to divided to tip, margins with distinct lobes or segments, apex acute to rounded, venation pinnate.

to 6 pairs, overlapping, strongly ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, asymmetrically cuneate, basal pinna pair often divided into 2 unequal parts, lobed to tip, margins entire, sinuate to shallowly crenate, apex rounded to truncate, venation pinnate.

Trophophore

stalk 0–1 mm;

blade dull to shiny green to dark green, deltate, 1–2-pinnate, to 6 × 7 cm.

stalk 0–10 mm, 0.1–0.5 times length of trophophore rachis;

blade dull, glaucous, whitish green, deltate, 2-pinnate, 4 × 6 cm, thickly leathery.

Sporophores

1–3-pinnate, 1–2.5 times length of trophophore, divided into several equally long branches (all other botrychiums have a single stalk or 1 dominant and 2 smaller).

1–3-pinnate, 1–1.5 times length of trophophore.

2n

=90.

=90.

Botrychium lanceolatum

Botrychium pumicola

Phenology Leaves appearing in summer.
Habitat Pumice scree
Elevation 1900–2500m (6200–8200ft)
Distribution
from FNA
AK; AZ; CO; CT; ID; KY; MA; ME; MI; MN; MT; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; ON; QC; SK; YT; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
OR
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium pumicola is a famous narrow endemic known from only a few colonies on fully exposed pumice scree on the sides of and in the general vicinity of Crater Lake, Klamath and Deschutes counties, Oregon. This plant has a very congested appearance with an extremely compact sporangial cluster and overlapping pinnae. Like most other members of subg. Botrychium, the trophophore is located high on the common stalk, but the common stalk is subterranean, giving the impression that the leaf originates near ground level. Botrychium pumicola has been found growing with B. lanceolatum and B. simplex.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trophophore blade green to pale yellow-green, broad, coarse, succulent; middle and terminal segments usually more than 2 mm wide exclusive of lobes.
subsp. lanceolatum
1. Trophophore blade dark green, slender, delicate but firm; middle segments usually less than 2 mm wide exclusive of lobes.
subsp. angustisegmentum
Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Subordinate taxa
B. lanceolatum subsp. angustisegmentum, B. lanceolatum subsp. lanceolatum
Synonyms Osmunda lanceolata
Name authority (S. G. Gmelin) Angström: Bot. Not. 1854: 68. (1854) Coville: in L. Underwood, Native Ferns ed. 6 69. (1900)
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