Botrychium lanceolatum
Botrychium lunarioides
botryche élancé, lance-leaf grapefern, lance-leaf moonwort, red triangle moonwort, triangle moonwort
prostrate grapefern, winter grapefern
to 5 pairs, ascending, approximate, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, linear to broadly lanceolate, entire to divided to tip, margins with distinct lobes or segments, apex acute to rounded, venation pinnate.
to 5 pairs, usually well separated, horizontal, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, divided to tip.
stalk 0–1 mm;
blade dull to shiny green to dark green, deltate, 1–2-pinnate, to 6 × 7 cm.
stalk 0.1–1 cm;
blades usually pale green, plane, 2–3-pinnate, to 8 × 12 cm, often much smaller, fleshy.
1–3-pinnate, 1–2.5 times length of trophophore, divided into several equally long branches (all other botrychiums have a single stalk or 1 dominant and 2 smaller).
2-pinnate, 1–2 times length of trophophores.
fan-shaped, margins denticulate, apex rounded, venation like ribs of fan, midrib absent.
=90.
=90.
Botrychium lanceolatum
Botrychium lunarioides
Subspecies 2 (2 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The stalk and proximal part of rachis of Botrychium lunarioides contains huge tracheidal idioblasts with annular thickenings, visible in cleared leaves (H.J. Arnott 1960). Another peculiarity of this species is the tendency for the sporophores to remain curled in late fall and early winter and to become erect in February. Botrychium lunarioides is often associated with Schizachyrium scoparius Michaux and Ophioglossum crotalophoroides Walter. The name B. biternatum was misapplied by L.Underwood to B. lunarioides.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Trophophore blade green to pale yellow-green, broad, coarse, succulent; middle and terminal segments usually more than 2 mm wide exclusive of lobes. | subsp. lanceolatum |
1. Trophophore blade dark green, slender, delicate but firm; middle segments usually less than 2 mm wide exclusive of lobes. | subsp. angustisegmentum |
- Local floras:
BC, 
OR, 
WA - Local Web sites:
CalFlora, CalPhotos, 
Flora NW, 
MI Flora, 
MN Wildflowers, 
PNW Herbaria, 
SW CO Wildflowers 
WildflowerSearch
iNaturalist (observations)
USDA Plants Database- LBJ Wildflower Center
- SEINet
- Plants of the World Online
- Encyclopedia of Life
- Wikipedia
- Google Image Search
LA Plants
