Botrychium lanceolatum
Botrychium echo
botryche élancé, lance-leaf grapefern, lance-leaf moonwort, red triangle moonwort, triangle moonwort
echo moonwort, reflected grape-fern
to 5 pairs, ascending, approximate, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, linear to broadly lanceolate, entire to divided to tip, margins with distinct lobes or segments, apex acute to rounded, venation pinnate.
to 4 pairs, spreading or only moderately ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, oblanceolate to linear-spatulate, ± parallel-sided, divided to tip, shallowly lobed or rarely 2-cleft, basal pinna cleft into single basiscopic projection and large acroscopic projection, margins entire, apex acute, venation pinnate.
stalk 0–1 mm;
blade dull to shiny green to dark green, deltate, 1–2-pinnate, to 6 × 7 cm.
stalk 0–4 mm;
blade shiny green, broadly oblong to oblong-deltate, 1–2-pinnate, to 4 × 3cm, firm.
1–3-pinnate, 1–2.5 times length of trophophore, divided into several equally long branches (all other botrychiums have a single stalk or 1 dominant and 2 smaller).
1–2 pinnate, 1–2 times length of trophophore rachis.
=90.
=180.
Botrychium lanceolatum
Botrychium echo
Subspecies 2 (2 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Botrychium echo is one of four moonwort species that commonly produce clusters of minute, spheric gemmae at the root bases. This species tends to have a reddish brown stripe along the common stalk from the base of the trophophore stalk.
Of conservation concern.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Trophophore blade green to pale yellow-green, broad, coarse, succulent; middle and terminal segments usually more than 2 mm wide exclusive of lobes. | subsp. lanceolatum |
1. Trophophore blade dark green, slender, delicate but firm; middle segments usually less than 2 mm wide exclusive of lobes. | subsp. angustisegmentum |
- Local floras:
BC, 
OR, 
WA - Local Web sites:
CalFlora, CalPhotos, 
Flora NW, 
MI Flora, 
MN Wildflowers, 
PNW Herbaria, 
SW CO Wildflowers 
WildflowerSearch
iNaturalist (observations)
USDA Plants Database- LBJ Wildflower Center
- SEINet
- Plants of the World Online
- Encyclopedia of Life
- Wikipedia
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