Botrychium lanceolatum |
Botrychium campestre |
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botryche élancé, lance-leaf grapefern, lance-leaf moonwort, red triangle moonwort, triangle moonwort |
botryche champètre, Iowa moonwort, prairie moonwort |
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Pinnae | to 5 pairs, ascending, approximate, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, linear to broadly lanceolate, entire to divided to tip, margins with distinct lobes or segments, apex acute to rounded, venation pinnate. |
to 5(–9) pairs, spreading, usually remote, separated 1–3 times pinna width, in some populations irregularly and extensively fused with considerable webbing along rachis, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to the adjacent pair, mostly linear to linear-spatulate, undivided to tip, margins crenulate to dentate, usually notched or cleft into 2 or several segments, apex rounded to acute, venation like ribs of fan, midrib absent. |
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Trophophore | stalk 0–1 mm; blade dull to shiny green to dark green, deltate, 1–2-pinnate, to 6 × 7 cm. |
stalk usually absent but sometimes broadly tapered to 10 mm in forms with coalesced proximal pinnae; blade glaucescent, oblong, longitudinally folded when alive, 1-pinnate, to 4 × 1.3 cm, very fleshy. |
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Sporophores | 1–3-pinnate, 1–2.5 times length of trophophore, divided into several equally long branches (all other botrychiums have a single stalk or 1 dominant and 2 smaller). |
1(–2, rarely)-pinnate, 1–1.5 times length of trophophore. |
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2n | =90. |
=90. |
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Botrychium lanceolatum |
Botrychium campestre |
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Phenology | Leaves appearing in early spring and dying in late spring and early summer, long before those of associated moonworts. | |||||
Habitat | Extremely inconspicuous in prairies, dunes, grassy railroad sidings, and fields over limestone | |||||
Elevation | 50–1200 m (200–3900 ft) | |||||
Distribution |
AK; AZ; CO; CT; ID; KY; MA; ME; MI; MN; MT; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; ON; QC; SK; YT; Eurasia
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CO; IA; MI; MN; MT; ND; NE; NY; SD; WI; WY; AB; ON; SK |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Botrychium campestre is one of four moonwort species that commonly produce dense clusters of minute, spheric gemmae at the root bases. Peculiar forms of B. campestre with coalescent pinnae are found on dunes in the vicinity of Lake Michigan. Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | Ophioglossaceae > Botrychium > subg. Botrychium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Osmunda lanceolata | |||||
Name authority | (S. G. Gmelin) Angström: Bot. Not. 1854: 68. (1854) | W. H. Wagner & Farrar: Amer. Fern J. 76: 39, figs. 2, 4, 5. (1986) | ||||
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