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botryche élancé, lance-leaf grapefern, lance-leaf moonwort, red triangle moonwort, triangle moonwort

northern moonwort

Pinnae

to 5 pairs, ascending, approximate, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, linear to broadly lanceolate, entire to divided to tip, margins with distinct lobes or segments, apex acute to rounded, venation pinnate.

to 6 pairs, ascending, mostly overlapping, distance between 1st and 2d pinnae only slightly greater than between 2d and 3d pairs, basal pinna pair usually considerably larger than adjacent pair, obliquely rhomboidal to oblanceolate-spatulate, mostly shallowly lobed to rarely pinnate, margins entire to very narrowly shallowly crenulate, apex pointed, venation pinnate only at bases of proximal pinnae, otherwise ± like ribs of fan.

Trophophore

stalk 0–1 mm;

blade dull to shiny green to dark green, deltate, 1–2-pinnate, to 6 × 7 cm.

stalk sessile or nearly so;

blade shiny green, ovate-deltate, 1–2-pinnate, to 6 cm, fleshy.

Sporophores

1–3-pinnate, 1–2.5 times length of trophophore, divided into several equally long branches (all other botrychiums have a single stalk or 1 dominant and 2 smaller).

1–2-pinnate, 1–1.5 times length of trophophore.

2n

=90.

Botrychium lanceolatum

Botrychium boreale

Phenology Leaves appearing in July and August.
Habitat Dry meadows, south-facing slopes
Elevation 200–600 m (700–2000 ft)
Distribution
from FNA
AK; AZ; CO; CT; ID; KY; MA; ME; MI; MN; MT; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; ON; QC; SK; YT; Eurasia
[WildflowerSearch map]
[BONAP county map]
from USDA
Greenland
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

This well-marked northern Eurasian species is best known in Scandinavia, where it occurs most commonly with Botrychium lunaria, with which it occasionally hybridizes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trophophore blade green to pale yellow-green, broad, coarse, succulent; middle and terminal segments usually more than 2 mm wide exclusive of lobes.
subsp. lanceolatum
1. Trophophore blade dark green, slender, delicate but firm; middle segments usually less than 2 mm wide exclusive of lobes.
subsp. angustisegmentum
Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Subordinate taxa
B. lanceolatum subsp. angustisegmentum, B. lanceolatum subsp. lanceolatum
Synonyms Osmunda lanceolata
Name authority (S. G. Gmelin) Angström: Bot. Not. 1854: 68. (1854) J. Milde: Bot. Zeitung 15(51): 880. (1857)
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