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botryche élancé, lance-leaf grapefern, lance-leaf moonwort, red triangle moonwort, triangle moonwort

point moonwort, tail grapefern

Pinnae

to 5 pairs, ascending, approximate, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, linear to broadly lanceolate, entire to divided to tip, margins with distinct lobes or segments, apex acute to rounded, venation pinnate.

to 6 pairs, spreading to ascending, mostly separated by pinna width, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to next adjacent pair, narrowly oblanceolate to linear-lanceolate, simple to irregularly divided to tip, margins commonly ± entire and with 1–several shallow and irregular lobes, apex acuminate, venation pinnate.

Trophophore

stalk 0–1 mm;

blade dull to shiny green to dark green, deltate, 1–2-pinnate, to 6 × 7 cm.

stalk 0–20 mm, 0.1–0.5 times length of trophophore rachis;

blade dull, glaucescent, ovate-oblong, 1-pinnate, to 6 × 5 cm, firm.

Sporophores

1–3-pinnate, 1–2.5 times length of trophophore, divided into several equally long branches (all other botrychiums have a single stalk or 1 dominant and 2 smaller).

1–2-pinnate, 1.4–2 times length of trophophore.

2n

=90.

=180.

Botrychium lanceolatum

Botrychium acuminatum

Phenology Leaves appearing in midspring, dying in fall.
Habitat Lake Superior region in sand dunes in shade, old fields, grassy railroad sidings, and roadside ditches
Elevation 200–500 m (700–1600 ft)
Distribution
from FNA
AK; AZ; CO; CT; ID; KY; MA; ME; MI; MN; MT; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; ON; QC; SK; YT; Eurasia
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[BONAP county map]
from FNA
MI; ON
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium acuminatum grows with various other moonworts, particularly its nearest relatives, B. hesperium and B. matricariifolium.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trophophore blade green to pale yellow-green, broad, coarse, succulent; middle and terminal segments usually more than 2 mm wide exclusive of lobes.
subsp. lanceolatum
1. Trophophore blade dark green, slender, delicate but firm; middle segments usually less than 2 mm wide exclusive of lobes.
subsp. angustisegmentum
Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Subordinate taxa
B. lanceolatum subsp. angustisegmentum, B. lanceolatum subsp. lanceolatum
Synonyms Osmunda lanceolata
Name authority (S. G. Gmelin) Angström: Bot. Not. 1854: 68. (1854) W. H. Wagner: Contr. Univ. Michigan Herb. 17: 321. (1990)
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