Botrychium hesperium |
Botrychium subg. Botrychium |
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western moonwort |
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Plants | less than 15 cm. |
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Roots | usually 10 or fewer, yellow or brown, 0.5–1.5mm diam. 1 cm from base. |
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Leaf | primordia glabrous.; leaf sheath closed. |
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Pinnae | to 6 pairs, ascending, usually approximate or overlapping except in shade forms, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair commonly much larger and more divided than adjacent pair, lobed to tip, basal pair oblong to oblong-lanceolate with lobed margins, remainder broadly spatulate with entire margins or 1 or more shallow lobes, apex rounded, venation pinnate. |
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Trophophore | stalk 0–3(–10) mm, to 1/4 length of trophophore rachis; blade ± gray-green, dull, oblong-linear to deltate, 1–2-pinnate, to 6 × 5 cm, firm. |
short-stalked or nearly sessile (long-stalked in forms of B. simplex and B. pedunculosum) arising from the middle or high on common stalk (low in some individuals of B. montanum, B. mormo, and B. simplex), blade usually 1 per plant, appearing in spring and dying in summer or fall, absent during winter, mostly linear to oblong to oblong deltate (deltate in B. lanceolatum), lobed to 1–2(–3)-pinnate, mostly less than 2.5 cm wide when mature, herbaceous to leathery. |
Sporophores | 1–3 pinnate, 2–3 times length of trophophore. |
long- or short-stalked, arising from middle to distal portion of common stalk, well above ground level (low on common stalk, near ground level in some forms of B. simplex), always present, stalks and rachis only slightly flattened, not fleshy, 0.5–2 mm wide. |
Common | stalk lacking idioblasts. |
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Pinna | lobes and segments, when present, asymmetric, either borne basiscopically or acroscopically. |
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x | =45. |
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2n | =180. |
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Botrychium hesperium |
Botrychium subg. Botrychium |
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Phenology | Leaves appearing in midspring, dying in early fall. | |
Habitat | Grassy mountain slopes, snow fields, road ditches with willows, and sand dunes | |
Elevation | 200–2800 m (700–9200 ft) | |
Distribution |
AZ; CO; ID; MI; MT; UT; WY; AB; BC; ON; SK
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Worldwide |
Discussion | In the Rocky Mountains Botrychium hesperium grows often with B. echo, and in the Lake Superior region, with B. acuminatum and B. matricariifolium. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Trophophore blades of most species in Botrychium subg. Botrychium are divided into lobes or segments and are not truly pinnate except in those species with fan-shaped segments. Very rarely, blades are more than 1-pinnate (western form of B. simplex). Species ca. 25 (21 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | Ophioglossaceae > Botrychium |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | B. matricariifolium subsp. hesperium | |
Name authority | (Maxon & R. T. Clausen) W. H. Wagner & Lellinger: Amer. Fern J. 71: 92. (1981) | Swartz |
Web links |