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western moonwort

botryche à segments spatulés, spathulate botrychium, spatulate moonwort, spoon-shape moonwort

Pinnae

to 6 pairs, ascending, usually approximate or overlapping except in shade forms, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair commonly much larger and more divided than adjacent pair, lobed to tip, basal pair oblong to oblong-lanceolate with lobed margins, remainder broadly spatulate with entire margins or 1 or more shallow lobes, apex rounded, venation pinnate.

to 8 pairs, ascending, remote, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, mostly narrowly spatulate to linear-spatulate and rounded or ± 2-cleft, lobed to unlobed to tip, margins mainly entire or occasionally irregularly and shallowly incised, apex rounded-notched, venation like ribs of fan, midrib absent.

Trophophore

stalk 0–3(–10) mm, to 1/4 length of trophophore rachis;

blade ± gray-green, dull, oblong-linear to deltate, 1–2-pinnate, to 6 × 5 cm, firm.

stalk 0–1 mm;

blade shiny yellow-green, narrowly deltate, flat, 1-pinnate, to 8 × 2.5 cm, thick, leathery.

Sporophores

1–3 pinnate, 2–3 times length of trophophore.

1–2-pinnate, 1.2–2 times length of trophophore.

2n

=180.

=180.

Botrychium hesperium

Botrychium spathulatum

Phenology Leaves appearing in midspring, dying in early fall. Leaves appearing late spring through summer.
Habitat Grassy mountain slopes, snow fields, road ditches with willows, and sand dunes Sand dunes, old fields, and grassy railroad sidings
Elevation 200–2800 m (700–9200 ft) 0–2000 m (0–6600 ft)
Distribution
from FNA
AZ; CO; ID; MI; MT; UT; WY; AB; BC; ON; SK
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[BONAP county map]
from FNA
AK; MI; MT; AB; BC; NB; NT; ON; PE; QC; YT
[BONAP county map]
Discussion

In the Rocky Mountains Botrychium hesperium grows often with B. echo, and in the Lake Superior region, with B. acuminatum and B. matricariifolium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium spathulatum has long been confused with the more common B. minganense, with which it often grows in the Lake Superior region. The leaves appear later in B. spathulatum than in B. minganense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. virginianum
Synonyms B. matricariifolium subsp. hesperium
Name authority (Maxon & R. T. Clausen) W. H. Wagner & Lellinger: Amer. Fern J. 71: 92. (1981) W. H. Wagner: Amer. Fern J. 80: 77. (1990)
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