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western moonwort

botryche à feuille de matricaire, chamomile grape-fern, daisy-leaf moonwort, daisyleaf grape-fern, matricary grape-fern

Pinnae

to 6 pairs, ascending, usually approximate or overlapping except in shade forms, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair commonly much larger and more divided than adjacent pair, lobed to tip, basal pair oblong to oblong-lanceolate with lobed margins, remainder broadly spatulate with entire margins or 1 or more shallow lobes, apex rounded, venation pinnate.

to 7 pairs, somewhat ascending, approximate to slightly remote, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pairs, spatulate-ovate to narrowly ovate, divided to tip, ultimate segments squarish to linear, rounded to square to acute at apex, margins entire to lobed to fully dissected, apex rounded to acute, venation pinnate.

Trophophore

stalk 0–3(–10) mm, to 1/4 length of trophophore rachis;

blade ± gray-green, dull, oblong-linear to deltate, 1–2-pinnate, to 6 × 5 cm, firm.

stalk 0–5 mm, to 1/6 length of trophophore rachis;

blade dull, ± glaucescent green, oblong, 1–3-pinnate, to 10 × 9 cm, firm.

Sporophores

1–3 pinnate, 2–3 times length of trophophore.

1–3-pinnate, 1.3–2.4 times length of trophophore.

2n

=180.

=180.

Botrychium hesperium

Botrychium matricariifolium

Phenology Leaves appearing in midspring, dying in early fall. Leaves appearing in spring, dying in late summer.
Habitat Grassy mountain slopes, snow fields, road ditches with willows, and sand dunes Old fields, secondary growth woods
Elevation 200–2800 m (700–9200 ft) 0–1200 m (0–3900 ft)
Distribution
from FNA
AZ; CO; ID; MI; MT; UT; WY; AB; BC; ON; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; DC; DE; IA; IL; IN; KY; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; NB; NF; NS; ON; QC; SPM; Europe
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[BONAP county map]
Discussion

In the Rocky Mountains Botrychium hesperium grows often with B. echo, and in the Lake Superior region, with B. acuminatum and B. matricariifolium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Synonyms B. matricariifolium subsp. hesperium B. lunaria var. matricariifolium
Name authority (Maxon & R. T. Clausen) W. H. Wagner & Lellinger: Amer. Fern J. 71: 92. (1981) (Döll) A. Braun ex W. D. J. Koch: Syn. Deut. Schweiz. Fl., ed. 2 7: 1009. (1847)
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