Botrychium hesperium |
Botrychium echo |
|
---|---|---|
western moonwort |
echo moonwort, reflected grape-fern |
|
Pinnae | to 6 pairs, ascending, usually approximate or overlapping except in shade forms, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair commonly much larger and more divided than adjacent pair, lobed to tip, basal pair oblong to oblong-lanceolate with lobed margins, remainder broadly spatulate with entire margins or 1 or more shallow lobes, apex rounded, venation pinnate. |
to 4 pairs, spreading or only moderately ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, oblanceolate to linear-spatulate, ± parallel-sided, divided to tip, shallowly lobed or rarely 2-cleft, basal pinna cleft into single basiscopic projection and large acroscopic projection, margins entire, apex acute, venation pinnate. |
Trophophore | stalk 0–3(–10) mm, to 1/4 length of trophophore rachis; blade ± gray-green, dull, oblong-linear to deltate, 1–2-pinnate, to 6 × 5 cm, firm. |
stalk 0–4 mm; blade shiny green, broadly oblong to oblong-deltate, 1–2-pinnate, to 4 × 3cm, firm. |
Sporophores | 1–3 pinnate, 2–3 times length of trophophore. |
1–2 pinnate, 1–2 times length of trophophore rachis. |
2n | =180. |
=180. |
Botrychium hesperium |
Botrychium echo |
|
Phenology | Leaves appearing in midspring, dying in early fall. | Leaves appearing in June, dying in September. |
Habitat | Grassy mountain slopes, snow fields, road ditches with willows, and sand dunes | Grassy mountain slopes, snow fields, road ditches, and sand dunes |
Elevation | 200–2800 m (700–9200 ft) | 2500–3700 m (8200–12100 ft) |
Distribution |
AZ; CO; ID; MI; MT; UT; WY; AB; BC; ON; SK
|
AZ; CO; UT
|
Discussion | In the Rocky Mountains Botrychium hesperium grows often with B. echo, and in the Lake Superior region, with B. acuminatum and B. matricariifolium. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Botrychium echo is one of four moonwort species that commonly produce clusters of minute, spheric gemmae at the root bases. This species tends to have a reddish brown stripe along the common stalk from the base of the trophophore stalk. Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | Ophioglossaceae > Botrychium > subg. Botrychium |
Sibling taxa | ||
Synonyms | B. matricariifolium subsp. hesperium | |
Name authority | (Maxon & R. T. Clausen) W. H. Wagner & Lellinger: Amer. Fern J. 71: 92. (1981) | W. H. Wagner: Amer. Fern J. 73: 57. (1983) |
Web links |