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crenulate moonwort, dainty moonwort, scalloped moonwort

Plants

less than 15 cm.

Roots

usually 10 or fewer, yellow or brown, 0.5–1.5mm diam. 1 cm from base.

Leaf

primordia glabrous.;

leaf sheath closed.

Pinnae

to 5 pairs, spreading, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, broadly fan-shaped, undivided to tip, margins mainly crenulate to dentate, proximal pinnae with 1 or more shallow incisions, apex rounded, apical lobe linear to linear-cuneate, well separated from adjacent lobes, venation like ribs of fan, midrib absent.

Trophophore

stalk 0.5–7 mm;

blade yellow-green, oblong, 1-pinnate, to 6 × 2 cm, thin, herbaceous.

short-stalked or nearly sessile (long-stalked in forms of B. simplex and B. pedunculosum) arising from the middle or high on common stalk (low in some individuals of B. montanum, B. mormo, and B. simplex), blade usually 1 per plant, appearing in spring and dying in summer or fall, absent during winter, mostly linear to oblong to oblong deltate (deltate in B. lanceolatum), lobed to 1–2(–3)-pinnate, mostly less than 2.5 cm wide when mature, herbaceous to leathery.

Sporophores

1–2-pinnate, 1.3–3 times length of trophophore.

long- or short-stalked, arising from middle to distal portion of common stalk, well above ground level (low on common stalk, near ground level in some forms of B. simplex), always present, stalks and rachis only slightly flattened, not fleshy, 0.5–2 mm wide.

Common

stalk lacking idioblasts.

Pinna

lobes and segments, when present, asymmetric, either borne basiscopically or acroscopically.

x

=45.

2n

=90.

Botrychium crenulatum

Botrychium subg. Botrychium

Phenology Leaves appearing in mid to late spring, dying in late summer; in extremely dry years of shorter duration or not appearing at all.
Habitat Local in marshy and springy areas
Elevation 1200–2500 m (3900–8200 ft)
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
Worldwide
Discussion

Botrychium crenulatum is commonly associated with B. simplex in California. In the Wallowa Mountains of Oregon it occurs with B. ascendens, B. lunaria, and B. minganense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Trophophore blades of most species in Botrychium subg. Botrychium are divided into lobes or segments and are not truly pinnate except in those species with fan-shaped segments. Very rarely, blades are more than 1-pinnate (western form of B. simplex).

Species ca. 25 (21 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Subordinate taxa
Name authority W. H. Wagner: Amer. Fern J. 71: 21. (1981) Swartz
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