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crenulate moonwort, dainty moonwort, scalloped moonwort

botryche à segments spatulés, spathulate botrychium, spatulate moonwort, spoon-shape moonwort

Pinnae

to 5 pairs, spreading, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, broadly fan-shaped, undivided to tip, margins mainly crenulate to dentate, proximal pinnae with 1 or more shallow incisions, apex rounded, apical lobe linear to linear-cuneate, well separated from adjacent lobes, venation like ribs of fan, midrib absent.

to 8 pairs, ascending, remote, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, mostly narrowly spatulate to linear-spatulate and rounded or ± 2-cleft, lobed to unlobed to tip, margins mainly entire or occasionally irregularly and shallowly incised, apex rounded-notched, venation like ribs of fan, midrib absent.

Trophophore

stalk 0.5–7 mm;

blade yellow-green, oblong, 1-pinnate, to 6 × 2 cm, thin, herbaceous.

stalk 0–1 mm;

blade shiny yellow-green, narrowly deltate, flat, 1-pinnate, to 8 × 2.5 cm, thick, leathery.

Sporophores

1–2-pinnate, 1.3–3 times length of trophophore.

1–2-pinnate, 1.2–2 times length of trophophore.

2n

=90.

=180.

Botrychium crenulatum

Botrychium spathulatum

Phenology Leaves appearing in mid to late spring, dying in late summer; in extremely dry years of shorter duration or not appearing at all. Leaves appearing late spring through summer.
Habitat Local in marshy and springy areas Sand dunes, old fields, and grassy railroad sidings
Elevation 1200–2500 m (3900–8200 ft) 0–2000 m (0–6600 ft)
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; MI; MT; AB; BC; NB; NT; ON; PE; QC; YT
[BONAP county map]
Discussion

Botrychium crenulatum is commonly associated with B. simplex in California. In the Wallowa Mountains of Oregon it occurs with B. ascendens, B. lunaria, and B. minganense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium spathulatum has long been confused with the more common B. minganense, with which it often grows in the Lake Superior region. The leaves appear later in B. spathulatum than in B. minganense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. virginianum
Name authority W. H. Wagner: Amer. Fern J. 71: 21. (1981) W. H. Wagner: Amer. Fern J. 80: 77. (1990)
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