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crenulate moonwort, dainty moonwort, scalloped moonwort

botryche à limbe rugueux, St. Lawrence grapefern, ternate grapefern

Pinnae

to 5 pairs, spreading, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, broadly fan-shaped, undivided to tip, margins mainly crenulate to dentate, proximal pinnae with 1 or more shallow incisions, apex rounded, apical lobe linear to linear-cuneate, well separated from adjacent lobes, venation like ribs of fan, midrib absent.

to 9 pairs, usually approximate, horizontal to ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, divided to tip.

Trophophore

stalk 0.5–7 mm;

blade yellow-green, oblong, 1-pinnate, to 6 × 2 cm, thin, herbaceous.

stalk 2 to 15 cm, 1–2.5 times length of trophophore rachis;

blade green, finely rugulose and convex distally, 2–4-pinnate, to 15 × 26 cm, somewhat herbaceous.

Sporophores

1–2-pinnate, 1.3–3 times length of trophophore.

2-pinnate, 1–2 times length of trophophore.

Pinnules

obliquely and angularly trowel-shaped to spatulate, margins usually denticulate, apex acute, venation pinnate.

2n

=90.

=90.

Botrychium crenulatum

Botrychium rugulosum

Phenology Leaves appearing in mid to late spring, dying in late summer; in extremely dry years of shorter duration or not appearing at all. Leaves green over winter, appearing in midspring.
Habitat Local in marshy and springy areas In open fields and secondary forests over wide range in vicinity of St. Lawrence Seaway
Elevation 1200–2500 m (3900–8200 ft) 200–1000 m (700–3300 ft)
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
MI; MN; NY; VT; WI; ON; QC
Discussion

Botrychium crenulatum is commonly associated with B. simplex in California. In the Wallowa Mountains of Oregon it occurs with B. ascendens, B. lunaria, and B. minganense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The name "rugulosum" refers to the tendency of the segments to become more or less wrinkled and convex. Botrychium rugulosum occurs with B. dissectum, B. multifidum, and rarely B. oneidense. It is often found in small stands of only 5–10 individuals, but some populations number over 100.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Sceptridium > sect. Sceptridium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. simplex, B. spathulatum, B. virginianum
Synonyms B. multifidum
Name authority W. H. Wagner: Amer. Fern J. 71: 21. (1981) W. H. Wagner: Contr. Univ. Michigan Herb. 15: 315. (1982)
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