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crenulate moonwort, dainty moonwort, scalloped moonwort

botryche élancé, lance-leaf grapefern, lance-leaf moonwort, red triangle moonwort, triangle moonwort

Pinnae

to 5 pairs, spreading, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, broadly fan-shaped, undivided to tip, margins mainly crenulate to dentate, proximal pinnae with 1 or more shallow incisions, apex rounded, apical lobe linear to linear-cuneate, well separated from adjacent lobes, venation like ribs of fan, midrib absent.

to 5 pairs, ascending, approximate, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, linear to broadly lanceolate, entire to divided to tip, margins with distinct lobes or segments, apex acute to rounded, venation pinnate.

Trophophore

stalk 0.5–7 mm;

blade yellow-green, oblong, 1-pinnate, to 6 × 2 cm, thin, herbaceous.

stalk 0–1 mm;

blade dull to shiny green to dark green, deltate, 1–2-pinnate, to 6 × 7 cm.

Sporophores

1–2-pinnate, 1.3–3 times length of trophophore.

1–3-pinnate, 1–2.5 times length of trophophore, divided into several equally long branches (all other botrychiums have a single stalk or 1 dominant and 2 smaller).

2n

=90.

=90.

Botrychium crenulatum

Botrychium lanceolatum

Phenology Leaves appearing in mid to late spring, dying in late summer; in extremely dry years of shorter duration or not appearing at all.
Habitat Local in marshy and springy areas
Elevation 1200–2500 m (3900–8200 ft)
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CO; CT; ID; KY; MA; ME; MI; MN; MT; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; ON; QC; SK; YT; Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

Botrychium crenulatum is commonly associated with B. simplex in California. In the Wallowa Mountains of Oregon it occurs with B. ascendens, B. lunaria, and B. minganense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trophophore blade green to pale yellow-green, broad, coarse, succulent; middle and terminal segments usually more than 2 mm wide exclusive of lobes.
subsp. lanceolatum
1. Trophophore blade dark green, slender, delicate but firm; middle segments usually less than 2 mm wide exclusive of lobes.
subsp. angustisegmentum
Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Subordinate taxa
B. lanceolatum subsp. angustisegmentum, B. lanceolatum subsp. lanceolatum
Synonyms Osmunda lanceolata
Name authority W. H. Wagner: Amer. Fern J. 71: 21. (1981) (S. G. Gmelin) Angström: Bot. Not. 1854: 68. (1854)
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